Petitina angulata ( Sieber, 1936 ) Harzhauser & Landau, 2012

Harzhauser, Mathias & Landau, Bernard, 2012, A revision of the Neogene Cancellariid Gastropods of the Paratethys Sea 3472, Zootaxa 3472, pp. 1-71 : 37

Petitina angulata ( Sieber, 1936) nov. comb.

Figs 8A 1–A 3, 8B 1–B 3

Cancellaria inermis Pusch — Franzl 1870: 49.

Cancellaria (Sveltia) inermis Pusch var. angulata n. v. Sieber 1936: 98, pl. 3, figs 7a–b [non Cancellaria angulata Eichwald, 1830 ].

Narona (Sveltia) inermis (Pusch, 1837) — Harzhauser 2002: 110 [non Petitina inermis (Pusch, 1837) ].

Type material. NHM Vienna ( Austria); see below for details .

Studied material. 3 spec. Niederkreuzstetten ( Austria) NHMW 1849 /0023/0016.

Illustrated material. Figs 8A 1–A 3: specimen illustrated in Sieber (1936, pl. 3, figs 7a–b), NHMW 1849 /0023/ 0016, Niederkreuzstetten ( Austria), height: 28.7 mm, width: 18.5 mm, syntype , Figs 8B 1–B 3: height: 24.8 mm, width: 14.8 mm, syntype.

Description: A medium sized cancellariid with gradate spire and elongated barrel-shaped last whorl. The spire whorls are high with nearly flat flanks, bulgy shoulder and narrow subsutural swelling. The high last whorl has subparallel flanks and a shoulder swelling that forms a narrow sutural ramp. A very weak concavity appears directly below the shoulder. The shell surface is smooth except for blunt growth lines and faint axial ribs on the early spire whorls. Base slowly contracting. A weak central columellar fold is adjoined by a faint anterior fold; columella straight. Columellar callus thickened grading into a prominent parietal callus. The outer lip is only fragmentarily preserved; no lirae or denticles are developed; no umbilicus.

Discussion. The thin outer lip, the moderately wide aperture with very prominent inner lip and the weak central columellar fold place the species in Petitina and points to a relation to Petitina inermis (Pusch, 1837) . Consequently, Sieber (1936) introduced the name angulata as variety name of Cancellaria inermis Pusch, 1837 for these specimens. This species, however, differs from Petitina inermis (Pusch, 1837) in its gradate spire, barrel-shaped last whorl, smaller size, the presence of a callus and the absence of axial sculpture, justifying a separation on species level. The name “ angulata ” is preoccupied by Cancellaria angulata Eichwald (1830) for a shell from Poland, which was later transferred to Turbinella by Eichwald (1853). As these taxa have not been considered to be congeneric after 1899, no replacement is needed in accordance with ICZN 23.9.5 and the valid cyronym is Petitina angulata ( Sieber, 1936) .

Distribution. This rare species is known so far only from the Karpatian (Late Burdigalian) of the Paratethys Sea.

Paratethys —Karpatian: Waschberg–Zdanice Unit ( Austria, Niederkreuzstetten).