Petitina inermis (Pusch, 1837) Harzhauser & Landau, 2012

Harzhauser, Mathias & Landau, Bernard, 2012, A revision of the Neogene Cancellariid Gastropods of the Paratethys Sea 3472, Zootaxa 3472, pp. 1-71 : 35-36

Petitina inermis (Pusch, 1837) nov. comb.

Figs 3L, 7H 1–H 3, 7I 1–I 2, 7J 1–J 3

Buccinum mitraeforme Pusch —Pusch in Andrzejowski 1830: 94, pl. 4, fig. 1.

Cancellaria inermis m.—Pusch 1837: 129, pl. 11, fig. 22.

Cancellaria inermis Pusch —Hörnes 1854: 313, pl. 34, figs 10–13.

Cancellaria inermis Pusch —Pereira da Costa 1867: 201, pl. 24, fig. 9, pl. 26, fig. 4.

Cancellaria inermis Pusch — Almera & Bofill 1884: 65, pl. F, figs 35–38.

Cancellaria inermis var. colligens Sacco, 1894: 22 .

Sveltia inermis Pusch — Friedberg 1914: 244, pl. 15, fig. 8, text–fig. 54.

Sveltia inermis (Pusch) mut. gallica nov. mut.— Peyrot 1928: 229, no. 1130, pl. 14, figs 11–14.

Sveltia inermis Pusch — Janoschek 1932: 76.

Cancellaria (Sveltia) inermis Pusch — Sieber 1936: 97.

Narona (Sveltia) inermis (Pusch) — Sieber 1936: 245.

Cancellaria (Sveltia) inermis Pusch — Csepreghy-Meznerics 1956: 416, pl. 9, figs 20–21.

Sveltia (Sveltia) cf. inermis (Pusch) — Erünal-Erentöz 1958: 89, pl. 13, figs 14–14a.

Cancellaria (Sveltia) inermis Pusch — Strausz 1966: 380, pl. 43, figs. 6–9.

Sveltia inermis (Pusch, 1837) — Zelinskaya et al. 1968: 217, pl. 49, figs 35–36.

Narona (Sveltia) inermis (Pusch, 1837) — Bałuk 1997: 47, pl. 16, figs 3–5.

Narona (Inermia) inermis (Pusch) — Schultz 1998: 72, pl. 29, fig. 6.

Narona inermis (Pusch, 1837) — Mikuž 2009: 29, pl. 9, figs 124–125.

non Narona (Sveltia) inermis (Pusch, 1837) — Harzhauser 2002: 110 [= Petitina angulata ( Sieber, 1936) ].

Type material. the collection of Pusch was destroyed during the Second World War ( Cleevely 1983)

Studied material. 32 spec. Grund ( Austria) NHMW 1855 / 0045/0733, 1851/0002/0052, NHMW A372 ; 2 spec. Niederleis ( Austria) NHMW 1860 /0015/0695; 5 spec. Guntersdorf ( Austria) NHMW A899 ; 17 spec. Enzesfeld ( Austria) NHMW 2009 z0098/0011, NHMW 2009 z0098/0012, NHMW 1846 / 0037/0295; 3. spec. Bad Vöslau ( Austria) NHMW 1852 / 0012/0037, NHMW 1874 /0024/0044; 3 spec. CoŞteiu de Sus ( Romania) NHMW 1867 / 0029/0127; NHMW 2012 /0004/0001; 3 spec. Lăpugiu de Sus ( Romania) NHMW 1873 /0025/0024; 3 spec. Korytnica ( Poland) NHMW 1853 /0038/0028.

Illustrated material. Fig. 3L: Petitina inermis (Pusch, 1837) , CoŞteiu de Sus ( Romania), height: 17.1 mm, width: 10.1 mm, NHMW 2012/0004/0001. Figs. 7J 1–J 3: specimen illustrated in Hörnes (1854, pl. 34, figs 10a–b), Grund ( Austria), height: 62.1 mm, width: 32.5 mm, NHMW 1855/0045/0733. Figs 7I 1–I 2: specimen illustrated in Hörnes (1854, pl. 34, figs 11a–b), Enzesfeld ( Austria), height: 44.4 mm, width: 23.7 mm, NHMW 2009z0098/ 0011. Figs 7H 1–H 3: specimen illustrated in Hörnes (1854, pl. 34, figs 12a–b), Enzesfeld ( Austria), height: 32.8 mm, width: 19.3 mm, NHMW 2009z0098/0012.

Discussion. This variable species comprises extraordinarily large specimens of more than 65 mm in height. Subadult shells are characterised by angulated spire whorls and a row of low nodes along the shoulder. Abapically, the nodes may grade into axial swellings or ribs. The angulation and the axial sculpture disappear on adult and large sized shells, which may form a sutural collar. The considerable morphologic variability contradicts a separation of the Viennese shells from those of the Polish–Carpathian Foredeep. This would be the case if the variety name colligens Sacco 1894 would be accepted for the shells illustrated in Hörnes (1854) as suggested by Sacco (1894). The protoconch is rarely preserved. Only a juvenile specimen from CoŞteiu de Sus in Romania is available with a protoconch of at least three whorls. Its initial part is low dome-shaped, with a strongly convex and rapidly expanding third whorl and incised sutures.

Buccinum mitraeforme Pusch in Andrzejowski, 1830 would have become a secondary homonym of Voluta mitraeformis Brocchi, 1814 in the first revision of this species by Pusch (1837). At that time both taxa were placed in Cancellaria . Therefore, Pusch (1837) introduced the replacement name Cancellaria inermis .

Unfortunately, Simionescu and Barbu (1940) used Buccinum mitraeforme again for a nassariid gastropod, which was later transferred to Duplicata Zhizhchenko in Kolesnikov, 1939 by Harzhauser and Kowalke (2004). This was also overlooked by Harzhauser and Cernohorsky (2011) in their nomenclatorial rectifications of nassariid gastropods. According to ICZN 23.9.5., however, a replacement name for this nassariid is not necessary as both taxa were not considered congeneric after 1899.

Distribution. This is a very common species in Badenian deposits of the Paratethys. During the Middle and Late Miocene it is also documented from the Atlantic coast (Pereira da Costa 1867; Peyrot 1928) and appears also in the Late Serravallian or Early Tortonian of the Turkish Karaman Basin (Erünal–Erentöz 1958) and the Late Miocene of Catalonia.

North East Atlantic —Serravallian: Aquitaine Basin ( France); Late Miocene: Adiça ( Portugal). Paratethys — Badenian: Polish–Carpathian Foredeep ( Poland: Korytnica); North Alpine Foreland Basin ( Austria: Grund, Windpassing, Guntersdorf); Vienna Basin ( Czech Republic: Hrušovany; Austria: Niederleis, Steinebrunn, Gainfarn, Enzesfeld, Gumpoldskirchen, Bad Vöslau, Baden–Sooss, Baden); Eisenstadt–Sopron Basin ( Austria: Forchtenau); Mecsek Mts. and Pannonian Basin ( Hungary: Szob, Hidas, Letkés); Transylvanian Basin ( Romania: Lăpugiu de Sus, CoŞteiu de Sus); Krka Basin ( Slovenia: Šentjernej, Gorenje Mokro Polje). Proto –Mediterranean Sea —Serravallian: Montjuic Basin ( Spain, Catalonia); Late Serravallian–Early Tortonian: Karaman Basin ( Turkey).