Chactopsoides gonzalezspongai, Ochoa & Rojas-Runjaic & Pinto-da-Rocha & Prendini, 2013

Ochoa, José A., Rojas-Runjaic, Fernando J. M., Pinto-da-Rocha, Ricardo & Prendini, Lorenzo, 2013, Systematic Revision Of The Neotropical Scorpion Genus Chactopsis Kraepelin, 1912 (Chactoidea: Chactidae), With Descriptions Of Two New Genera And Four New Species, Bulletin of the American Museum of Natural History 2013 (378), pp. 1-121 : 97-105

Chactopsoides gonzalezspongai , n. sp.

Figures 2 , 4B , 5D , 9D , 45C, D , 46C, D , 48A , 49A , 50C , 51B , 52C, D , 58–61 ; table 5

TYPE MATERIAL: Holotype ³ ( MHNLS 1424 ), 6 ³, 5 ♀ paratypes ( MHNLS), 4 ³,

3 ♀, 3 juv. paratypes ( AMNH), 2 juv. paratypes ( AMNH [ LP 10093, 10094]), 1 ³, 1 ♀, 3 juv. paratypes ( MHNC), VENEZUELA: Amazonas: Municipio Atabapo: Isla Piñate, confluence of Orinoco and Ventuari rivers, 03 ° 58912.480 N 67 ° 01949.860W, 92 m, 9.viii. 2009, F. Rojas-Runjaic, A. Ferrer, and J.A. Ochoa, forest.

ETYMOLOGY: The specific name is a patronym honoring the late Manuel Angel González-Sponga (1929–2009) for his contributions to the knowledge of the biodiversity of Venezuelan arachnids.

DIAGNOSIS: Chactopsoides gonzalezspongai , n. sp., may be separated from C. anduzei , n. comb., by the hemispermatophore morphology: the lamina is approximately the same length as the trunk in C. gonzalezspongai , n. sp., but 15 % –20 % shorter in C. anduzei , n. comb.; the median lobe and dorsal apophysis are less developed in C. gonzalezspongai , n. sp., than C. anduzei , n. comb. (figs. 57, 61); the sheath-shaped part occupies two-thirds of the trunk in C. gonzalezspongai , n. sp. (fig. 61), compared with 80 % of trunk in C. anduzei , n. comb. (fig. 57). Other differences between the two species are as follows: the pedipalp chela manus length/width ratio is 4.00–4.58 (³) and 3.82–4.15 (♀) in C. gonzalezspongai , n. sp., compared with 4.43–4.82 (³) and 4.21– 4.67 (♀) in C. anduzei , n. comb.; the LIM carinae of metasomal segments II and III are absent in the anterior third and represented by only a few granules posteriorly in C. gonzalezspongai , n. sp. (fig. 48A), whereas they are restricted to the anterior third of segments II and III in C. anduzei , n. comb. (fig. 47B); metasomal segments II–IV each exhibit three complete stripes of pigmentation (single VM and paired VL), and the VM stripe is complete in C. gonzalezspongai , n. sp., compared with two complete stripes of pigmentation (VL only), whereas the VM stripe is restricted to the anterior half of each segment in C. anduzei , n. comb. Chactopsoides gonzalezspongai , n. sp., differs from C. marahuacaensis , n. comb., as follows. The carapace is coarsely to finely granular, especially anteriorly and laterally (more so in ³), in C. gonzalezspongai , n. sp. (fig. 45C, D), but mostly smooth, with scattered granules near anterior margin only, and nongranular surfaces punctate, in C. marahuacaensis , n. comb. (fig. 45E). The metasomal VSM carinae are complete and granular on segments II–IV in C. gonzalezspongai , n. sp. (fig. 49A), whereas the VSM carinae are obsolete on segment II, and complete and granular on segments III and IV in C. marahuacaensis , n. comb. (fig. 49B). The DMA carina of the pedipalp chela is costate on the distal quarter of the manus and on the fixed finger in C. gonzalezspongai , n. sp., but costate on the fixed finger only in C. marahuacaensis , n. comb. Chelal trichobothrium db is situated equidistant between esb and eb in C. gonzalezspongai , n. sp., but closer to eb in C. marahuacaensis , n. comb. The dimensions of metasomal segment V and telson provide additional differences between C. gonzalezspongai , n. sp., C. anduzei , n. comb., and C. marahuacaensis , n. comb. The length/width ratio of metasomal segment V is 1.63–1.90 (³) and 1.63–1.72 (♀) in C. gonzalezspongai , n. sp., 1.83–2.10 (³) and 1.75–1.96 (♀) in C. anduzei , n. comb., and 1.77–1.82 (♀) in C. marahuacaensis , n. comb. The telson length/height ratio is 2.72–3.28 (³) and 2.63–3.13 (♀) in C. gonzalezspongai , n. sp., 3.22–3.75 (³) and 3.40–3.85 (♀) in C. anduzei , n. comb., and 3.13–3.15 (♀) in C. marahuacaensis , n. comb.

DESCRIPTION: Based on the holotype ³ and paratypes. Measurements of the holotype ³ and a paratype ♀ recorded in table 5 .

Total length: ³, 21.00– 30.7 mm (n 5 11; mean 5 26.01 mm); ♀, 30.39–33.30 (n 5 8; mean 5 31.8 mm).

Color: Base color brown to brownish yellow (adults; fig. 5D) or yellow (juveniles); carapace, tergites, sternite VII, metasoma, legs, and pedipalps with dark brown spots; sternites III–VI and coxosternal region yellow; pectines white; aculeus dark brown. Cheliceral manus, dorsal surfaces with fine reticulate pigmentation, becoming contiguous distally near base of fixed finger; fixed and movable fingers entirely pigmented. Carapace, median ocular tubercle and surfaces around lateral ocelli very densely pigmented; lateral and posterolateral surfaces densely pigmented, anteromedian longitudinal, postocular, and posteromedian longitudinal sulci unpigmented. Pedipalp femur, dorsal and external surfaces densely pigment- ed with small, unpigmented areas medially, ventral and internal surfaces faintly pigment- ed, with pigmentation stripes along DI, DE, VI, and EM carinae; patella, dorsal surface with irregular pigmentation and pigmentation stripes along DE and DI carinae, external surface with pigmentation stripe along EM carina and additional reticulate pigmentation, ventral surface with pigmentation stripes along VI and VE carinae, pigmentation on internal surface concordant with granulation; chela with pigmentation stripes along D, SD, DS, DMA, DI, E, IM, VI, and VE carinae, contiguous at base of fingers, VI and VE pigmentation stripes faint or absent. Legs, femur, prolateral surfaces faintly pigmented; patella and tibia pro- and retrolateral surfaces faintly pigmented. Tergites densely pigmented except for small, unpigmented area medially, forming narrow median stripe longitudinally across tergites, restricted to anterior half of tergite VII in some adults. Coxosternal region, genital operculum and pectines mostly unpigmented. Sternites III–VI, mostly unpigmented, except for faint spots anterolaterally; VII with two broad, dark pigmentation stripes sublaterally and faint pigmentation submedially. Metasomal segments I–IV, dorsal surfaces each with pigmentation stripes along DSM and DL carinae, surfaces between DSM carinae with conspicuous subtriangular spots of pigmentation, sometimes divided by narrow unpigmented stripe; lateral surfaces each with pigmentation stripes along ML carinae and reticulate pigmentation along LIM and LSM carinae, surfaces between DL and ML carinae densely pigmented in posterior third of segment, surfaces between ML and VL carinae densely pigmented in posterior twothirds; ventral surfaces each with paired pigmentation stripes along VL carinae, surfaces between VL and VSM carinae densely pigmented in posterior two-thirds of segment I, posterior half of II and III, and posterior third of IV, surfaces between VSM carinae densely pigmented, forming narrow stripe, not contiguous with lateral pigmentation posteriorly. Metasomal segment V, dorsal surface with DSM pigmentation in anterior half of segment and pigmentation stripes along DL carinae; lateral surface with reticulate pigmentation along LIM and LSM carinae, and dense pigmentation stripes along ML carinae, becoming contiguous with DL pigmentation posteriorly, surfaces between ML and VL carinae densely pigmented in posterior third of segment; ventral surface with three dense pigmentation stripes along VM and paired VL carinae, contiguous in posterior third of segment, and with fine pigmentation stripes along VSM carinae in anterior third, surfaces between VL and VSM carinae densely pigmented in posterior two-thirds. Telson vesicle, dorsolateral margins with faint pigmentation in anterior half of segment; lateral and ventral surfaces more densely pigmented in juveniles; ventral surface with two broad VL and one narrow VM pigmentation stripes, separated by two narrow, unpigmented stripes.

Chelicerae: Movable finger with well-developed serrula, occupying slightly more than half its length; ventral subdistal tooth present.

Carapace: Anterior margin with shallow, biconcave median notch and several microsetae (fig. 45C, D); posterior margin sublinear, with shallow median notch and few microsetae. Surfaces entirely covered with variable fine and medium granulation (³, fig. 45C), becoming more fine posterolaterally, or granulation restricted to interocular, circumocular, anterolateral and median lateral surfaces (♀, fig. 45D), granulation more pronounced in ³; nongranular surfaces punctate, more so in ♀; scattered microsetae throughout. Median ocelli half an ocular diameter apart. Anteromedian longitudinal sulcus finely granular, nongranular surfaces punctate; posteromedian longitudinal sulcus punctate; other sulci smooth.

Pedipalps: Femur, length/width ratio, ³, 2.84–3.13 (n 5 11; mean 5 3.02); ♀, 2.77– 3.16 (n 5 8, mean 5 2.95); DE, DI, and VI carinae complete, granular (fig. 59A); EM carina restricted to distal third of segment, weakly granular (³) or smooth (♀); VM carina granular, restricted to proximal third of segment; VE carina vestigial, reduced to three or four granules proximally; IM carina vestigial, reduced to three prominent, isolated granules in proximal half of segment; dorsal intercarinal surface with variable fine and coarse granulation, more dense in ³, and slightly punctate in ♀, and with few coarse granules medially; external intercarinal surface almost smooth, punctate distally; ventral intercarinal surface smooth and punctate; internal intercarinal surface finely granular (³) or punctate (♀). Patella, length/width ratio, ³, 2.63–2.95 (n 5 11; mean 5 2.77); ♀, 2.45–2.61 (n 5 8, mean 5 2.54); DE, DI, and VI carinae complete, granular, DE slightly less developed (fig. 59B–D); EM and VE carinae obsolete, reduced to punctation and, in VE carina of some ³, to weak granulation in proximal third in ³; DPP comprising moderate proximal granule and additional smaller granules, VPP reduced to one or two smaller granules; dorsal intercarinal surface densely granular, more so in ³, nongranular surfaces punctate, more so in ♀; external and ventral intercarinal surfaces smooth and punctate; internal intercarinal surface densely granular. Chela manus narrow (³) or slightly incrassate (♀), fingers relatively elongated (fig. 60); chela length/width ratio, ³, 4.00–4.58 (n 5 11; mean 5 4.31); ♀, 3.82–4.15 (n 5 8, mean 5 3.97); length/height ratio, ³, 4.00–4.70 (n 5 11; mean 5 4.34); ♀, 3.93–4.15 (n 5 8, mean 5 3.99). Manus and fingers, intercarinal surfaces weakly granular, nongranular surfaces densely punctate, and covered with scattered microsetae; D carina discontinuous, punctate along entire length, becoming weakly granular in distal third of manus and costate on distal two-thirds of fixed finger; SD carina restricted to proximal third of manus, punctate, and with two or three granules proximally; DS carina complete, weakly granular and punctate on manus, becoming costate on fixed finger; DMA carina weakly granular and punctate in proximal threequarters of manus, becoming costate in distal quarter and on fixed finger, well developed on distal quarter of manus and proximal third of fixed finger, becoming progressively less pronounced distally on finger until trichobothrium dm 2; DI carina discontinuous, interrupted by two porous areas at base of fixed finger, weakly granular and punctate on manus, costate on distal three-quarters of fixed finger; E, VE, and VI carinae punctate; IM carina weakly granular and punctate.

Fixed finger, median denticle row continuous, complete; flanked by nine external and eight internal denticles; internal denticles not interspersed with accessory denticles; internal accessory denticles arranged in one discontinuous row, comprising 21 small denticles; external accessory denticles arranged in one continuous row, interspersed with external denticles.

Trichobothria: Femur with three trichobothria (fig. 59A). Patella with 33 trichobothria (fig. 59B–D): two dorsal, seven ventral, 23 external, one internal; trichobothria v 6 and v 7 situated submedially, v 6 equidistant between v 5 and v 7; est 5 situated on VE margin and slightly distal to est 4; est 2 situated proximal to other est trichobothria, est 3 situated in same axis as est 4; em 1 – em 3 usually situated slightly proximal to or in same axis as em 2; esb 2 situated distal to esb 3. Chela with 26 trichobothria (fig. 60): 10 situated on manus, three ventral, seven external; 16 situated on fixed finger, seven external, six dorsal, three internal (it, ist, ib); isb absent; it situated between et 3 and est; Est situated closer to Et 1 than to V 3; Et 2 situated distal to Et 1; Esb situated proximal to Eb 2; eb situated near base of fixed finger; db situated between esb and eb; dm 1 situated slightly distal to et 3.

Legs: Prolateral surfaces granular, retrolateral surfaces smooth. Femur III, DI carina restricted to distal half of segment; DE carina obscured by dorsal granulation; EM and VI carinae complete. Patella III, DI carina complete; VI carina comprising few granules, restricted to distal third of segment; other carinae obscured by dorsal and external granulation. Basitarsus III setose; two dorsal rows of small subspiniform granules, retrodorsal row on proximal two-thirds of segment, prodorsal row restricted to medial third; one dorsal and two ventral rows of small brushlike spinules, retrodorsal and proventral rows restricted to distal third of segment, ventromedian row restricted to distal two-thirds. Telotarsus III setose, pro- and retroventral rows each with 5–7 elongated macrosetae.

Tergites: Pretergites I–VII, surfaces punctate. Posttergites I–VI, surfaces finely granular in anterior half, coarsely and densely granular in posterior half, more so on III–VI (³) or punctate and sparsely granular along posterior margin of I–III and sparsely granular in posterior half of IV–VI (♀); dorsomedian and dorsosubmedian carinae vestigial in posterior half of III–VI (³) or IV–VI (♀). Posttergite VII, surface coarsely granular (more so in ³); paired dorsosubmedian and dorsolateral carinae well developed in posterior two-thirds; posterior margin with transverse row of coarse granules.

Sternum: Ventral surface punctate, with six macrosetae (in one case eight), one pair situated anteriorly and two pairs on posterolateral lobes; apex and anterior margins with several microsetae (fig. 46C, D).

Pectines: Pectinal tooth count: ³, 10 (n 5 23), 11 (7); ♀, 7 (9), 8 (17) (fig. 46C, D).

Sternites: Sternites III–VI, surfaces smooth, punctate; VII, surface smooth and punctate with scattered granules along lateral margins, VL carinae vestigial, each comprising two granules in medial third.

Metasoma: Segments I– IV, dorsal intercarinal surfaces finely and densely granular (³) or smooth, punctate (♀) (figs. 48A, 49A) ; lateral and ventral intercarinal surfaces coarsely and densely granular (more so in ³), nongranular surfaces punctate (more so in ♀) (fig. 49A); small porous area situated posteriorly at LIM position on segments I– IV, slightly more developed on I than on II– IV; DSM carinae moderately developed, complete, granular on segment I, vestigial in anterior two-thirds of segments II–IV (less evident, obscured by fine granulation in ³); DL carinae complete, granular, posterior granules forming low mound, more pronounced on segments III and IV; LSM carinae vestigial, weakly evident, comprising few small granules in medial third (♀) or obscured by granulation (³) ; ML carinae complete, granular on segments I–IV, posterior granules slightly larger than others, forming low mound, on I–III; LIM carinae complete on segment I, reduced to few granules posteriorly on II–IV; VL carinae complete, granular, slightly more developed on segment IV; VSM carinae absent or obsolete, comprising two or three granules, on segment I, complete on II – IV, though less developed on II than on III and IV (fig. 49A). Segment V, length/width ratio: ³, 1.63–1.90 (n 5 11; mean 5 1.78), ♀, 1.63–1.72 (n 5 8, mean 5 1.67) ; length/height ratio, ³, 1.96–2.24 (n 5 11; mean 5 2.08), ♀, 1.85–2.03 (n 5 8, mean 5 1.95); dorsal intercarinal surface finely granular, nongranular surfaces punctate; lateral and ventral intercarinal surfaces densely granular, more coarsely so in ♀; small porous area situated posteriorly at LIM position; DL and VL carinae complete, granular (more developed in ♀) ; LSM carinae vestigial, weakly evident in anterior two-thirds of segment (♀), or obscured by granulation (³) ; ML carinae restricted to anterior twothirds of segment; LIM carinae absent; VM carina coarsely granular, restricted to anterior two-thirds of segment, obscured by scattered granulation in posterior third; VSM carinae absent (³) or vestigial, comprising row of small granules restricted to anterior third (♀) (fig. 51B) .

Telson: Length/height ratio, ³, 2.72–3.53 (n 5 11, mean 5 2.98); ♀, 2.63–3.13 (n 5 8, mean 5 2.97). Vesicle globose, more so in ³; dorsal intercarinal surface smooth; lateral and ventral intercarinal surfaces finely and densely granular (³) or coarsely granular, nongranular surfaces punctate (♀) (fig. 52C, D); ventral surface with several scattered microsetae. Aculeus short and gently curved (³) or slightly elongated and more strongly curved (♀).

Hemispermatophore: Lamina narrow, slen- der, approximately same length as or slightly shorter than trunk (fig. 61A–C); apex slender, relatively elongated, curved, and tapering distally; flagellum slightly undulated, approximately one-third the length of lamina; ental fold absent; articular flexure present, weakly developed; slight proximal constriction, forming elongated pedicel. Trunk well developed, elongated; sheath-shaped part well developed, occupying two-thirds of trunk, with median longitudinal sulcus; ventral concavity reduced to distal third; foot well developed, extending almost entire length of trunk. Lobe region reduced, with single lobe; ental lobe absent; median lobe reduced, extending to ventral surface, finely and sparsely papillose entally, without spines or papillae distally; median trough deep, extending entire length of median lobe; dorsal apophysis well developed, sclerotized along margin, laminar, slightly folded toward dorsal surface.

DISTRIBUTION: Chactopsis gonzalezspongai , n. sp., is known only from the type locality in the state of Amazonas, Venezuela (fig. 2).

HABITAT: Specimens were collected by UV light detection at night, from leaf litter in primary rainforest (fig. 4B), on a small island at the confluence of the Orinoco and Ventuari rivers.

Figures

Fig. 2. Distribution of Chactopsoides, n. gen., and Megachactops, n. gen., plotting known locality records in Brazil, Colombia, and Venezuela. Chactopsoides anduzei (González-Sponga, 1982), n. comb., solid squares; Chactopsoides gonzalezspongai, n. sp., solid diamond; Chactopsoides marahuacaensis (González-Sponga, 2004), n. comb., open square; Chactopsoides yanomami (Lourenço et al., 2011), n. comb., solid circle; Megachactops coriaceo (González-Sponga, 1991), n. comb., open diamond; Megachactops kuemoi, n. sp., open circle; undetermined Chactopsoides species, solid stars.

Fig. 4. Chactopsoides, n. gen., and Megachactops, n. gen., habitat. A. Autana, Amazonas, Venezuela, habitat of Chactopsoides anduzei (González-Sponga, 1982), n. comb., and Megachactops kuemoi, n. sp. B. Isla Piñate, confluence of Orinoco and Ventuari rivers, Amazonas, Venezuela, habitat of Chactopsoides gonzalezspongai, n. sp.

Fig. 5. Chactopsis Kraepelin, 1912, Chactopsoides, n. gen., and Megachactops, n. gen., habitus in life. A, B. Chactopsis amazonica Lourenço and Francke, 1986, Reserva Ducke, Manaus, Amazonas, Brazil. A. ³. B. ♀. C. Chactopsoides anduzei (González-Sponga, 1982), n. comb., ³, Isla Ratón, Amazonas, Venezuela. D. Chactopsoides gonzalezspongai, n. sp., ³, Isla Piñate, Amazonas, Venezuela. E. Chactopsis insignis Kraepelin, 1912, ³, Tahuayo River, Loreto Department, Peru. F. Megachactops kuemoi, n. sp., ♀, Tobogan del Cuao, Amazonas, Venezuela.

Fig. 9. Chactopsis Kraepelin, 1912, Chactopsoides, n. gen., and Megachactops, n. gen., dextral pedipalp chela, movable finger, dorsal aspect showing dentition. A. Chactopsis curupira, n. sp., paratype ³ (MZSP). B. Chactopsis insignis Kraepelin, 1912, ³ (MHNC). C. Chactopsoides anduzei (González-Sponga, 1982), n. comb., ³ (AMNH). D. Chactopsoides gonzalezspongai, n. sp., paratype ³ (AMNH). E. Megachactops kuemoi, n. sp., paratype ♀ (MHNC). Scale bars 5 1 mm.

Fig. 45. Chactopsoides, n. gen., carapace, dorsal aspect. A, B. Chactopsoides anduzei (González- Sponga, 1982), n. comb. A. ³ (AMNH). B. ♀ (AMNH). C, D. Chactopsoides gonzalezspongai, n. sp. C. Paratype ³ (AMNH). D. Paratype ♀ (AMNH). E. Chactopsoides marahuacaensis (González-Sponga, 2004), n. comb., holotype ♀ (MAGS). Scale bars 5 1 mm.

Fig. 46. Chactopsoides, n. gen., sternum and pectines, ventral aspect. A, B. Chactopsoides anduzei (González-Sponga, 1982), n. comb. A. ³ (AMNH). B. ♀ (AMNH). C, D. Chactopsoides gonzalezspongai, n. sp. C. Paratype ³ (AMNH). D. Paratype ♀ (AMNH). E. Chactopsoides marahuacaensis (González- Sponga, 2004), n. comb., holotype ♀ (MAGS). Scale bars 5 1 mm.

Fig. 48. Chactopsoides, n. gen., metasomal segments III–V, lateral aspect. A. Chactopsoides gonzalezspongai, n. sp., paratype ³ (AMNH). B. Chactopsoides marahuacaensis (González-Sponga, 2004), n. comb., holotype ♀ (MAGS). Scale bars 5 1 mm.

Fig. 49. Chactopsoides, n. gen., metasomal segments I–IV, ventral aspect. A. Chactopsoides gonzalezspongai, n. sp, paratype ³ (AMNH). B. Chactopsoides marahuacaensis (González-Sponga, 2004), n. comb., holotype ♀ (MAGS). Scale bars 5 1 mm.

Fig. 50. Chactopsoides, n. gen., metasomal segment V, lateral aspect. A, B. Chactopsoides anduzei (González-Sponga, 1982), n. comb. A. ³ (AMNH). B. ♀ (AMNH). C. Chactopsoides gonzalezspongai, n. sp., paratype ♀ (AMNH). Scale bars 5 1 mm.

Fig. 51. Chactopsoides, n. gen., metasomal segment V, ventral aspect. A. Chactopsoides anduzei (González-Sponga, 1982), n. comb., ³ (AMNH). B. Chactopsoides gonzalezspongai, n. sp., paratype ♀ (AMNH). C. Chactopsoides marahuacaensis (González-Sponga, 2004), n. comb., holotype ♀ (MAGS). Scale bars 5 1 mm.

Fig. 52. Chactopsoides, n. gen., telson, lateral aspect. A, B. Chactopsoides anduzei (González-Sponga, 1982), n. comb. A. ³ (AMNH). B. ♀ (AMNH). C, D. Chactopsoides gonzalezspongai, n. sp. C. Paratype ³ (AMNH). D. Paratype ♀ (AMNH). E. Chactopsoides marahuacaensis (González-Sponga, 2004), n. comb., holotype ♀ (MAGS). Scale bars 5 1 mm.

Fig. 58. Chactopsoides gonzalezspongai, n. sp., habitus. A, B. Paratype ³ (MHNC). C, D. Paratype ♀ (MHNC) A, C. Dorsal aspect. B, D. Ventral aspect. Scale bar 5 10 mm.

Fig. 59. Chactopsoides gonzalezspongai, n. sp., paratype ³ (AMNH), dextral pedipalp segments. A. Femur. B–D. Patella. A, B. Dorsal aspect. C. External aspect. D. Ventral aspect. Scale bar 5 1 mm.

Fig. 60. Chactopsoides gonzalezspongai, n. sp., dextral pedipalp chela. A. Paratype ³ (AMNH). B–E. Paratype ♀ (AMNH). A, C. External aspect. B. Dorsal aspect. D. Ventral aspect. E. Internal aspect. Scale bars 5 1 mm.

Fig. 61. Chactopsoides gonzalezspongai, n. sp., paratype ³ (AMNH), sinistral hemispermatophore. A. Ental aspect. B. Dorsal aspect. C. Ectal aspect. Scale bar 5 1 mm.