Chileanthicus, Werner, 1966

Kejval, Zbyněk, 2009, Taxonomic revision of the genus Chileanthicus Werner (Coleoptera: Anthicidae) 2180, Zootaxa 2180 (1), pp. 1-82 : 6-9

publication ID

https://doi.org/ 10.11646/zootaxa.2180.1.1

persistent identifier

https://treatment.plazi.org/id/3949251E-FFA8-776F-FF63-DDBBFA02FB03

treatment provided by

Felipe

scientific name

Chileanthicus
status

 

Key to the species of Chileanthicus (except C. lafertei )

1 Elytra with pale transverse band in posterior half; subapical process of metafemora minute; male sternum VII more or less distinctly emarginate postero-medially, with peg-like setae; species occuring in Chile................................. 2

- Elytra with different colour marking, never with single transverse band in posterior half; subapical process of metafemora usually more prominent; male sternum VII may be slightly modified, but never emarginate posteromedially; species occuring in Madagascar or Australia.............................................................................................. 9

2 Apical portion of tegmen of aedeagus with paired latero-basal incision ( Fig. 52 View FIGURES 45–53 ); female tergum VII strongly bulging ( Fig. 53 View FIGURES 45–53 ) ............................................................................................................................................... C. penai Werner

- Apical portion of tegmen lacking latero-basal incisions; female tergum VII at most moderately convex ................ 3

3 Elytra in female tapering apically ( Fig. 240 View FIGURES 232–241 ); apex of tegmen of aedeagus upturned in lateral view ( Fig. 12 View FIGURES 9–16 ) ............ .................................................................................................................................................. C. acutipennis sp. nov.

- Elytra in female evenly rounded to truncate apically; apex of tegmen straight to moderately curved ventrad ......... 4

4 Antennae conspicuously slender, slightly enlarged in terminal third; antennomere III more than 3.0 times as long as wide ( Fig. 32 View FIGURES 27–33 ) .............................................................................................................................................................. 5

- Antennae somewhat stronger, moderately enlarged in distal third, antennomere III less than 2.5 times as long as wide ( Fig. 33 View FIGURES 27–33 ) .............................................................................................................................................................. 6

5 Paired prongs of male sternite VIII slender, each with more than 10 peg-like setae, rather unevenly scattered ventrally ( Figs 18, 19 View FIGURES 17–26 ); apical portion of tegmen of aedeagus rather straight in lateral view, and apex wider, rounded in ventral view ( Fig. 20 View FIGURES 17–26 ) .................................................................................................................. C. elmorado sp. nov.

- Paired prongs of male sternite VIII more robust, each with less than 10 peg-like setae arranged in row ventrally ( Figs 28, 29 View FIGURES 27–33 ); apical portion of tegmen of aedeagus arcuate in lateral view, and apex narrowed, nearly bluntly pointed in ventral view ( Fig. 31 View FIGURES 27–33 ) ..................................................................................................................... C. hirsutus sp. nov.

6 Male sternum VII with about 40 peg-like setae ( Fig. 13 View FIGURES 9–16 ); apical portion of tegmen of aedeagus with conspicuous, long, transverse edge dorsally in basal half, distinctly protruding in lateral view ( Fig. 16 View FIGURES 9–16 ) ..... C. baculentus sp. nov.

- Male sternum VII with less than 20 peg-like setae; apical portion of tegmen rather smoothly shaped dorsally, dorsal edge indistinct or at least less conspicuous, short, barely protruding from outline of tegmen in lateral view ........... 7

7 Paired prongs of male sternite VIII with peg-like setae arranged somewhat laterally in row ( Figs 46, 47 View FIGURES 45–53 ); dorsal outline of apical portion of tegmen distinctly sinuous in lateral view, apex of tegmen rather wide, subtriangular in ventral view ( Fig. 48 View FIGURES 45–53 ) ............................................................................................................................... C. mitis sp. nov.

- Paired prongs of male sternite VIII with peg-like setae rather scattered on ventral side; dorsal outline of apical portion of tegmen at most slightly sinuous/impressed ( Fig. 26 View FIGURES 17–26 ) or evenly arcuate ( Fig. 41 View FIGURES 34–44 ) in lateral view, apex of tegmen rather narrowed, bluntly pointed in ventral view (only C. maritimus sp. nov., Fig. 42 View FIGURES 34–44 ) .................................... 8

8 Male sternum VII moderately and narrowly emarginate posteriorly ( Fig. 22 View FIGURES 17–26 ); apical portion of tegmen of aedeagus with dorsal transverse edge distant from its base ( Fig. 26 View FIGURES 17–26 , marked by arrow); female sternum VII modified, with pair of short, bluntly pointed median protrusions ( Fig. 21 View FIGURES 17–26 ) ..................................................... C. femineus sp. nov. (male)

- Male sternum VIII rather strongly and widely emarginate posteriorly ( Fig. 38 View FIGURES 34–44 ); apical portion of tegmen of aedeagus with dorsal transverse edge situated near base ( Fig. 41 View FIGURES 34–44 ); female sternum VII simple, with posterior margin evenly rounded ...................................................................................................................................... C. maritimus sp. nov.

9 Species occuring in Madagascar............................................................................................................................... 10

- Species occuring in Australia................................................................................................................................... 11

10 Body surface matte, very densely and finely punctured ( Fig. 58 View FIGURES 54–62 ), including lateral sides of pronotum; body setation very short, appressed, erect setae less conspicuous; paired prongs of male sternite VIII bearing peg-like setae ( Fig. 61 View FIGURES 54–62 ) .................................................................................................................................................. C. dentatus (Dajoz)

- Body surface moderately glossy, its dorsal punctation coarser and sparser ( Fig. 54 View FIGURES 54–62 ), lateral sides of pronotum partly impunctate and glossy; body setation generally longer and more raised, erect setae more conspicuous; paired prongs of male sternite VIII at most finely setose, lacking peg-like setae ( Fig. 56 View FIGURES 54–62 ) ................................... C. cursor sp. nov.

11 Metafemora unidentate; with single tooth-like process on inner side subapically ................................................... 12

- Metafemora bidentate or tridentate, with 1–2 processes on inner side and the third one on outer side subapically 41

12 Pronotum with dorso-lateral margins forming sharp edge ( Fig. 241 View FIGURES 232–241 ; see also C. comes sp. nov. with nearly carinate margins) .................................................................................................................................................................... 13

- Pronotum more or less rounded dorso-laterally (at most with short edge, close behind the widest point of pronotum, see C. comes sp. nov.) ............................................................................................................................................... 16

13 Head with tempora forming blunt edge at level of ventral eye margin (eyes thus seem to be situated rather dorso-laterally; Fig. 241 View FIGURES 232–241 ); punctation of elytra less conspicuous, simple, punctures distinctly smaller/finer than those on pronotum ........................................................................................................................................................................ 14

- Head with tempora rounded (eyes situated laterally); punctation of elytra conspicuous, its coarse punctures about as large as those on pronotum ....................................................................................................................................... 15

14 Head moderately glossy, distinctly but somewhat sparsely punctured, punctures finer/smaller than those on pronotum ( Fig. 102 View FIGURES 102–111 ); male characters as in Figs 104–107 View FIGURES 102–111 ......................................................................... C. discus sp. nov.

- Head matte, distinctly and very densely punctured, similar to pronotum ( Fig. 108 View FIGURES 102–111 ); male characters as in Figs 110, 111 View FIGURES 102–111 ........................................................................................................................................................ C. exul sp. nov.

15 Head widely rounded posteriorly in dorsal view ( Fig. 63 View FIGURES 63–71 ); antennae rather short, antennomeres less elongate ( Fig. 64 View FIGURES 63–71 ): III 1.8 times, X 1.2 times, XI 1.9 times as long as wide; male characters as in Figs 66, 67 View FIGURES 63–71 .................................. .................................................................................................................................................. C. angulicollis sp. nov.

- Head angulate posteriorly in dorsal view ( Fig. 129 View FIGURES 123–134 ); antennae longer, antennomeres more elongate ( Fig. 130 View FIGURES 123–134 ): III 2.3 times, X 1.7 times, XI 2.1 times as long as wide; male characters as in Figs 132–134 View FIGURES 123–134 ................. C. inflatus sp. nov.

16 Basal punctation of elytra distinctly double, with coarse and fine punctures ........................................................... 17

- Basal punctation of elytra uniformly fine to somewhat heterogeneous (indistinctly double, its coarse punctures barely discernible) ..................................................................................................................................................... 24

17 Erect setae of elytra conspicuous, much longer than ordinary, subdecumbent setae ............................................... 18

- Erect setae of elytra inconspicuous, about as long as other setae ............................................................................. 20

18 Paler marking and silvery setae of elytra forming narrow band/spot in apical and irregular spot in basal half, extending towards base ( Figs 146 View FIGURES 146–154 , 235 View FIGURES 232–241 ), silvery setae less contrasting, rather evenly directed posteriad (not swirled); subapical process of metafemora smaller ( Fig. 147 View FIGURES 146–154 ); male characters as in Figs 148, 149 View FIGURES 146–154 .............. C. kinchega sp. nov.

- Paler marking and silvery setae of elytra forming two, narrow transverse bands, silvery setae rather conspicuous, distinctly swirled ( Figs 86 View FIGURES 81–89 , 141 View FIGURES 135–145 ); subapical process of metafemora somewhat more conspicuous, with wider base ( Figs 87 View FIGURES 81–89 , 142 View FIGURES 135–145 ) ............................................................................................................................................................ 19

19 Male characters as in Figs 143–145 View FIGURES 135–145 .............................................................................................. C. interruptus (Lea)

- Male characters as in Figs 88, 89 View FIGURES 81–89 ................................................................................................ C. decipiens sp. nov.

20 Head and pronotum wide, head angulate posteriorly in dorsal view; basal coarse punctures of elytra distinctly larger than fine punctures ( Fig. 155 View FIGURES 155–159 ); male metatibiae with two longitudinal grooves on inner side, another male characters as in Figs 157–159 View FIGURES 155–159 ........................................................................................................................... C. latibasis (Lea)

- Head and pronotum narrower, head at most widely rounded posteriorly; basal coarse punctures of elytra at most slightly to moderately larger than fine punctures ( Figs 77 View FIGURES 72–80 , 123 View FIGURES 123–134 , 135 View FIGURES 135–145 , 171 View FIGURES 171–182 ); male matatibiae lacking grooves on inner side ............................................................................................................................................................................ 21

21 Elytra slightly glossy to matte, very densely punctured ........................................................................................... 22

- Elytra moderately glossy, somewhat less densely punctured ................................................................................... 23

22 Dorsal punctation of head distinctly double, coarse punctures much larger than fine punctures ( Fig. 123 View FIGURES 123–134 ); the anterior setose band of elytra extending dorso-laterally towards base ( Figs 123 View FIGURES 123–134 , 234 View FIGURES 232–241 ); male characters as in Figs 125–128 View FIGURES 123–134 ................................................................................................................................................... C. improvidus sp. nov.

- Dorsal punctation of head indistinctly double, coarse punctures only moderately larger than fine punctures ( Fig. 77 View FIGURES 72–80 ); the anterior setose band narrow, at most moderately widened near lateral margin; male characters as in Figs 79, 80 View FIGURES 72–80 . ........................................................................................................................................................... C. comes sp. nov.

23 Male sternum III with small median protuberance, male metatibiae with small process/denticle on inner side apically ( Fig. 173 View FIGURES 171–182 ), other male characters as in Figs 174–176 View FIGURES 171–182 ................................................................... C. melasomus (Lea)

- Male sternum III simple, lacking median protuberance, male metatibiae simple, lacking apical process/denticle, other male characters as in Figs 137–140 View FIGURES 135–145 ...................................................................................... C. insignis sp. nov.

24 Body largely rufous, elytra with black marking ( Fig. 166 View FIGURES 160–170 ), evenly and sparsely punctured, rather glossy, uniformly setose; male characters as in Figs 168–170 View FIGURES 160–170 ............................................................................... C. matthewsi sp. nov.

- Body largely dark coloured, elytra dark, unicoloured or with paler marking, densely punctured, at most moderately glossy, with silvery setose bands/spots ..................................................................................................................... 25

25 Elytra uniformly dark, basal punctation of elytra always uniformly fine ................................................................. 26

- Elytra mostly dark with distinct paler marking (bands/spots, suture, lateral margins), rarely nearly unicoloured (variability, some specimens of C. melasomus and C. quadrimaculatus ); basal punctation of elytra uniformly fine to somewhat heterogeneous (indistinctly double, its coarse punctures barely discernible, see C. melasomus ) ........... 32

26 Silvery/whitish setose bands of elytra with setae at least partly distinctly obliquely to transversely pointing (swirled) ................................................................................................................................................................................... 27

- Silvery/whitish setose bands of elytra with setae rather evenly pointing posteriad as ordinary appressed/subdecumbent setae ( Fig. 238 View FIGURES 232–241 ) .................................................................................................................................................. 31

27 Head large, distinctly wider (behind eyes) than narrow pronotum ( Fig. 90 View FIGURES 90–101 ); elytra rather short and ovoid, as long as head and pronotum ( Figs 90 View FIGURES 90–101 , 232 View FIGURES 232–241 ); pronotal disc with moderate median longitudinal impression; femora (especially metafemora) long clavate, rather abruptly swollen distally ( Fig. 93 View FIGURES 90–101 ) ............................................... C. audax sp. nov.

- Head smaller in relation to rather wide pronotum; elytra more elongate, longer than head and pronotum; pronotal disc evenly convex, median longitudinal impression at most indicated anteriorly; femora clavate, but their basal narrow portion rather evenly widening ( Figs 186, 189 View FIGURES 183–191 ) ................................................................................................ 28

28 Pronotum nearly trapezoidal, widely rounded to angulate anteriorly in dorsal view, strongly narrowing posteriad ( Fig. 183 View FIGURES 183–191 ); punctation of head and pronotum simple, punctures much larger than those on elytra; basal punctures of elytra arranged in short, obligue rows, separated by fine wrinkles ( Fig. 183 View FIGURES 183–191 )........................... C. pubifasciatus (Lea)

- Pronotum more or less widely rounded anteriorly and moderately narrowing posteriad in dorsal view ( Figs 72 View FIGURES 72–80 , 118 View FIGURES 112–122 , 185 View FIGURES 183–191 ); punctures of head and pronotum at most moderately larger than those on elytra ( Fig. 185 View FIGURES 183–191 ), if some of them distinctly larger, than punctation of head and pronotum distinctly double ( Figs 72 View FIGURES 72–80 , 118 View FIGURES 112–122 ); basal punctures of elytra irregularly scattered, interspaces rather smooth, unwrinkled ........................................................................................... 29

29 Elytra subtruncate apically; surface of head, pronotum and elytra nearly matte, very densely punctured ( Fig. 185 View FIGURES 183–191 ); silvery/whitish setae of transverse bands of elytra mostly transversely pointing laterad ( Figs 185 View FIGURES 183–191 , 237 View FIGURES 232–241 ) ..................... ........................................................................................................................................................ C. pulcher sp. nov.

- Elytra conjountly rounded apically; surface of head, pronotum and elytra rather moderately glossy, its punctation somewhat less dense ( Figs 72 View FIGURES 72–80 , 118 View FIGURES 112–122 ); silvery/whitish setae of transverse bands on elytra differently arranged, swirled medially ( Figs 72 View FIGURES 72–80 , 118 View FIGURES 112–122 ) ............................................................................................................................................. 30

30 Male characters as in Figs 120–122 View FIGURES 112–122 .............................................................................................. C. gammon sp. nov.

- Male characters as in Figs 74–76 View FIGURES 72–80 ............................................................................................... C. campicola sp. nov.

31 Antennae moderately enlarged in terminal third: antennomere X 1.7 times, XI 2.2 times as long as wide; pronotum trapezoidal, widely rounded to angulate anteriorly in dorsal view, strongly narrowing posteriad ( Fig. 183 View FIGURES 183–191 ); punctures of head and pronotum much larger than those on elytra, basal punctures of elytra arranged in short, obligue rows, which may appear to be separated by fine wrinkles ( Fig. 183 View FIGURES 183–191 ) ................................................ C. pubifasciatus (Lea)

- Antennae distinctly enlarged in terminal third ( Fig. 188 View FIGURES 183–191 ): antennomere X as long as wide, XI 1.8 times as long as wide; pronotum widely rounded anteriorly in dorsal view, moderately narrowing posteriad ( Fig. 187 View FIGURES 183–191 ); punctures of head and pronotum only moderately larger than those on elytra, basal punctures of elytra irregularly scattered ( Fig. 187 View FIGURES 183–191 ) ................................................................................................................................................ C. pumilio sp. nov.

32 Silvery setae of elytra both forming setose bands/spots (on paler marking) and scattered on dark coloured surface ( Fig. 233 View FIGURES 232–241 ) ................................................................................................................................................................... 33

- Silvery setae of elytra restricted to paler coloured marking and forming only setose bands/spots (they may be scattered at most along lateral margins, but never on dorsal side) .................................................................................. 34

33 Antero-lateral outline of pronotum evenly rounded in dorsal view ( Fig. 81 View FIGURES 81–89 ); male meso- and metatibiae slightly angulately produced on inner side apically, metatibiae uniformly short setose; male sternum III with small median protuberance, another male characters as in Figs 83–85 View FIGURES 81–89 ........................................................... C. communis sp. nov.

- Antero-lateral outline of pronotum with slight angle at widest place in dorsal view ( Fig. 112 View FIGURES 112–122 ); male meso- and metatibiae simple apically, metatibiae with a tuft of long setae on inner side apically ( Fig. 115 View FIGURES 112–122 ); male sternum III simple, male characters as in Figs 116–117 View FIGURES 112–122 ............................................................................................... C. eyrensis sp. nov.

34 Dorsal surface of head matte, very densely, evenly punctured, punctures nearly contiguous and uniform ( Fig. 204 View FIGURES 201–208 ); sternum VII with a pair of distinct protuberances ( Fig. 206 View FIGURES 201–208 ), another male characters as in Figs 207, 208 View FIGURES 201–208 ................. .................................................................................................................................................. C. tuberculifer sp. nov.

- Dorsal surface of head slightly glossy, rarely nearly matte ( C. comes sp. n.), punctures distinctly separated, often unevenly spaced (sparser in median row) and heterogeneous (more or less distinctly double); male sternum VII simple, lacking paired protuberance ............................................................................................................................... 35

35 Head widely rounded to angulate posteriorly; elytra somewhat widely rounded apically ( Fig. 68 View FIGURES 63–71 ); elytra nearly matte, very densely punctured, basal punctation simple, punctures somewhat serial (surface appears to be transversely wrinkled); anterior silvery setose band of elytra wide, its posterior margin uneven, and setae along anterior margin pointing mediad ( Fig. 68 View FIGURES 63–71 ); male characters as in Figs 70, 71 View FIGURES 63–71 ............................................ C. aprilis sp. nov.

- Head evenly to moderately widely rounded posteriorly; elytra evenly rounded apically; elytra less densely punctured, rarely nearly matte ( C. comes sp. n.), basal punctation simple or somewhat heterogeneous (indistinctly double), punctures rather evenly spaced (interspaces smooth, unwrikled); anterior silvery setose band of elytra rather narrow and even, setae along its anterior margin pointing posteriad ........................................................................ 36

36 Basal punctation of elytra simple, uniformly fine ( Figs 177 View FIGURES 171–182 , 192 View FIGURES 192–200 , 212 View FIGURES 209–219 ); male sternum VII simple, lacking median protuberance .............................................................................................................................................................. 37

- Basal punctation of elytra indistinctly double (its coarse punctures sometimes barely discernible, Figs 135 View FIGURES 135–145 , 171 View FIGURES 171–182 ); male sternum VII simple or with median protuberance ............................................................................................ 39

37 Head and pronotum wider, head rather widely rounded posteriorly ( Fig. 177 View FIGURES 171–182 ); dorsal punctation of head very dense, indistinctly double ( Fig. 177 View FIGURES 171–182 ); male metatibiae simple and with a tuft of long, stiff setae apically ( Fig. 179 View FIGURES 171–182 ), another male characters as in Figs 180–182 View FIGURES 171–182 .............................................................................................. C. paganus sp. nov.

- Head and pronotum narrower, head narrower, at most unevenly rounded posteriorly ( Figs 192 View FIGURES 192–200 , 212 View FIGURES 209–219 ); dorsal punctation of head less dense, clearly double, its coarse punctures distinctly larger then fine punctures .......................... 38

38 Head evenly densely punctured, including frons medially; male characters as in Figs 194–196 View FIGURES 192–200 ................................. ............................................................................................................................................ C. quadrimaculatus (King)

- Head unevenly punctured, punctures distinctly sparser medially, with indication of glossy medial line, especially anteriorly on frons and clypeus; male characters as in Figs 214–216 View FIGURES 209–219 ......................................... C. volselifer sp. nov.

39 Elytra slightly glossy to matte, punctures densely spaced; pronotum distinctly wider than head ( Fig. 77 View FIGURES 72–80 ), pronotal disc with nearly carinate lateral margins; male sternum III with median protuberance, male metatibiae uniformly short setose; male characters as in Figs 79–80 View FIGURES 72–80 ................................................................................. C. comes sp. nov.

- Elytra moderately glossy, punctures less densely spaced; pronotum narrower, pronotal disc more convex, with rounded lateral margins; male sternum VII simple or with median protuberance .................................................... 40

40 Male sternum III with small median protuberance, male metatibiae with small apical process/denticle ( Fig. 173 View FIGURES 171–182 ), another male characters as in Figs 174–176 View FIGURES 171–182 ................................................................................. C. melasomus (Lea)

- Male sternum III simple, lacking median protuberance, male metatibiae simple apically, lacking apical process, another male characters as in Figs 137, 140 View FIGURES 135–145 .................................................................................. C. insignis sp. nov.

41 Metafemora bidentate, with single process on inner side, which is strong, tooth-like, wide basally and strongly narrowing towards pointed apex ( Figs 151 View FIGURES 146–154 , 201 View FIGURES 201–208 ) ........................................................................................................... 42

- Metafemora bi- to tridentate, 1–2 processes on inner side narrow, thorn-like shaped ( Figs 95 View FIGURES 90–101 , 161 View FIGURES 160–170 ), the smaller one may be reduced ( Fig. 190 View FIGURES 183–191 ) or resembling angulately produced edge ( Fig. 211 View FIGURES 209–219 ) ...................................................... 43

42 Body darker coloured, blackish; pronotal disc uneven, with distinct median longitudinal impression; surface of elytra uneven, distinctly impressed dorsally at place of basal setose band; silvery setose bands of elytra very conspicuous, contrasting, setae largely and quite distinctly swirled ( Fig. 239 View FIGURES 232–241 ); body punctation conspicuously double; male characters as in Figs 198–200 View FIGURES 192–200 ...................................................................................................... C. speciosus (King)

- Body paler coloured, mostly brownish; pronotal disc and elytra evenly shaped, lacking distinct impressions; silvery setose bands of elytra less conspicuous, narrower, setae moderately swirled medially ( Fig. 150 View FIGURES 146–154 ); body punctation distinctly double but coarse punctures comparatively less conspicuous ( Fig. 150 View FIGURES 146–154 ); male characters as in Figs 153, 154 View FIGURES 146–154 ................................................................................................................................................. C. kingii (MacLeay)

43 Head angulate posteriorly; elytra with single silvery/whitish setose band ( Figs 209 View FIGURES 209–219 , 225 View FIGURES 220–231 ), erect setae rather short and inconspicuous; terminal spurs of metatibiae well developed ........................................................ C. uhmanni sp. nov.

- Head at most widely rounded posteriorly; elytra with two silvery/whitish setose bands ( Figs 94 View FIGURES 90–101 , 160 View FIGURES 160–170 , 236 View FIGURES 232–241 ), erect setae rather long and conspicuous; terminal spurs of metatibiae absent or at least strongly reduced and indistinct ..... ................................................................................................................................................................................... 44

44 Setation of metatibiae rather evenly short and fine (except apical fringe of short, stiff setae); male characters as in Figs 162–165 View FIGURES 160–170 .................................................................................................................................. C. mastersii (King)

- Setation of metatibiae with short, stiff, bluntly pointed setae scattered on outer side (except apical fringe) and numerous conspicuously long, raised setae on inner side distally ( Figs 96, 97 View FIGURES 90–101 ); male characters as in Figs 98–101 View FIGURES 90–101 ... ....................................................................................................................................................... C. dentivarius (Lea)

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Anthicidae

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