Pupina pallens Moellendorff , 1894

Jirapatrasilp, Parin, Sutcharit, Chirasak & Panha, Somsak, 2022, Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia, ZooKeys 1119, pp. 1-115 : 1

Pupina pallens Moellendorff, 1894

Figs 21N, O , 24B, C

Pupina pallens Möllendorff, 1894: 155, pl. 16, figs 27, 28. Type locality: Samui Islands, Gulf of Siam [Samui Island, Surat Thani Province, Thailand]. Fischer and Dautzenberg 1904: 431. BEDO 2017: 92. Sutcharit et al. 2018: fig. 5-13i.

Pupina (Tylotoechus) pallens - Kobelt 1902: 318, 319. Laidlaw 1928: 34. Zilch 1957: 47, pl. 2, fig. 16. Hemmen and Hemmen 2001: 39.

Type material examined.

Lectotype SMF 109951 (Figs 21N , 24B ) and paralectotypes SMF 109952 (4 shells), SMF 109953 (2 shells) from Golf von Siam: Koh Samui.

Other material examined.

CUMZ 12041 (1 shell) from Bang Phu Temple, Sam Roi Yot District, Prachuap Khiri Khan Province, 19 Oct. 2020. CUMZ 12042 (14 shells; Figs 21O , 24C ) from Suan Wiwek Bureau of Monks, Sam Roi Yot District, Prachuap Khiri Khan Province, 21 Oct. 2020.

Diagnosis.

Shell ovate; last whorl ca. three quarters of shell height. Apertural lip slightly thickened, not expanded; basal position widened. Both parietal and columellar teeth fin-shaped and slightly thickened; parietal tooth covering posterior canal; columellar tooth next to slit-like anterior canal.

Differential diagnosis.

Pupina pallens can be distinguished from all other species in the P. artata species group from mainland Southeast Asia by the widened basal position of the apertural lip.

Distribution.

The type locality ( Laidlaw 1928) and Prachuap Khiri Khan Province, western Thailand.

Remarks.

von Möllendorff (1894) stated that this species is different from P. arula in having "the more obtuse spire, the more distorted last whorl, and consequently the aperture placed more to the right and protracted at the base, the thinner outer peristome, the broader columella, the broad triangular parietal lamella, and the narrower lower incision". More sampling of this species, with both morphometric and molecular phylogenetic analyses, are needed to resolve the relationship between P. pallens and other species in the P. artata species group.

Figures

Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T).

Figure 24. A Pupina artata, specimen CUMZ 12038 from Tham Suea Temple, Krabi B, C Pupina pallens B lectotype SMF 109951 and C specimen CUMZ 12042 from Suan Wiwek Bureau of Monks, Prachuap Khiri Khan D-F Pupina limitanea D, E syntypes NHMUK 1903.7.1.2967 from East of Burma & Siam and F specimen CUMZ 12044 from Mae Lana junction, Mae Hong Son G Pupina bensoni sp. nov., holotype CUMZ 12045 / 1. Photo: H. Taylor, NHM (D, E).