Diochus hanagarthi, Irmler, 2017

Irmler, Ulrich, 2017, A review of the Neotropical genus Diochus ERICHSON, 1840 (Coleoptera: Staphylinidae: Staphylininae), Beiträge Zur Entomologie = Contributions to Entomology 67 (1), pp. 1-62 : 10-11

Diochus hanagarthi spec. nov.

urn:lsid:zoobank.org:act:00554886-6F3A-4FBA-9639-39AF0EE7E01B

Figs 9A–C , 38G , 41H

Type material: male Holotype: Peru: Huanuco, Llullapichis, Panguana station (74°56'W, 9°37'S), cut forest planted with Manihot, 6.10.1975, leg. W. Hanagarth ( UIC).

Paratypes: Ecuador: Pastaza Prov., 1000 m elev., 25 km N Puyo, 1000 m elev., Berlese , cloud forest litter, female, 13.7.1976, leg. S. Peck ( FMNH) ; Peru: same place as holotype, but Cocha , 4 females, 9.5.1976, leg. W. Hanagarth ( UIC) ; same place as holotype, cut forest, 3 males, 2 females, 4.5.1976, leg. W. Hanagarth ( UIC) ; same place as holotype, pitfall trap, river bank pasture, 10.– 24.1.1984, leg. M. Verhaagh ( UIC) ; Dept. Loreto, Campamento San Jacinto (75°51.77'W, 2°18.75'S), 175–215 m elevation, collected by flight intercept trap, female, 3.7.1993, leg. R. Leschen, #10 ( KNHM) ; Loreto Dept. Iquitos , 90 m elevation, collected by light intercept trap, female, 5.5.1992, leaf litter in the forest, female, 8.5.1992, leg. J. Danoff-Berg ( KNHM) ; Tambopata Prov. , 15 km NE Pto. Maldonado (69°03'W, 12°33'S), 200 m elevation, under leaves by stream, 1 male, 4 females, 5.7.1989, 1 male, from liana fruit fall, 25.6.1989 leg. J. Ashe, R. Leschen #397 #269 ( KNHM) ; Junin Dept., La Merced- Villa Rica Rd. (10°57.12'S, 75°17.30'W), 900 m elev., flight intercept trap, female, 15.– 21.10.1999, leg. R. Brooks, # PERU 1B99090 ( KNHM) ; Cuzco Dept., Consuelo, Manu rd. km 165, litter along stream, 1 male, 1 female, 4.10.1982, leg. L.E. Watrous & G. Mazurek ( FMNH) ; Madre do Dios Dept., Tambopata , litter at swamp edge, 2 females, 28.10.1982, leg. L.E. Watrous & G. Mazurek ( FMNH) ; Bolivia: Santa Cruz Dept., 3.7 km SSE Buena Vista, Hotel Flora y Fauna (63°33.15'W, 17°29.95'S), 400–440 m elevation, primary forest, flight intercept trap, female, 3.– 9.11.2002, leg. R. Leschen #BOL1L02- 052 ( KNHM) ; Cochabamba, 117 km E Yungas, Lagunitas (17°06.2'S, 65°40.57'W), 1000 m elev., montane forest litter, female, 1.2.1999, leg. R. Anderson #BOLA99014 ( KNHM) ; Cochabamba, 67.5 km NE Est. Biol. Valle del Sajita, Univ. de San Simon (17°06.33'S, 64°47.52'W), rain forest litter, female, 7.2.1999, leg. R. Anderson, #BOL1A99027 ( KNHM) ; Vaca Diez Prov., Beni Dept. , 2 km NW Tumichucua, forest, leaf litter in sunken area, female, 12. 8.1990, leg. P. Parrillo ( FMNH) ; Brazil: Amazonas, Manaus , Ilha de Marchanteria (59°58'W, 3°15'S), floating meadow, 2 males, 29.4.1981, leg. J. Adis ( UIC) ; Paraguay : without more data, coll Kraatz ( SDEI) ; Paraguari, Sapucai (25°40'S, 56°55'W), 190 m elev., light trap, 1 male, 3 females, 4.– 8.2.1995, leg. U. Drechsel ( KNHM) ; San Pedro, Vaca Jhu (24°34'S, 56°37'W), 120 m elev., 1 male, 3 females, 5.– 8.3.1993. leg. Drechsel ( KNHM) ; Guairá, Calle Florida , at light, 2 females, 7.2.1994, leg. Drechsel ( KNHM) ; same Prov., Melgarejo, Tacuara Creek , flood detritus, female, 20.10.1994, leg. Drechsel ( KNHM) ; Alto Paraguay, Madrejon , at light, 2 males, 1 female, 15.5.1995, leg. Drechsel (2 KNHM, 1 UIC) ; Argentina: Catamarca, Cencepción , female, 5.12.1971, leg. L. Herman ( AMNH) ; San Juan Prov., S. of Astica Chucuma (31°04'S, 67°17'W), leaf litter nr. stream, female, 4.11.1995, leg. L. Herman ( AMNH) ; San Juan Prov., NW of San Augustin de Valle Fertil, La Rinconada (30°47'S, 67°28'W), leaf litter nr. stream, 1100 m elev., female, 5.11.1995, leg. L. Herman ( AMNH) ; Tucuman, 15 km N Tucuman, Rio Sali , female, 30.12.1971, leg. L. Herman ( AMNH) ; San Luis Prov., Aguas Buenas (32°35.05'S, 64°57.01'W), litter nr. stream, female, 9.12.2000, leg. L. Herman ( AMNH) .

Diagnosis: D. hanagarthi closely resembles D. inornatus in size, colouration and by the thick and long seminal vesical of the endophallus. It can be separated from D. inornatus by the shorter head and temples. EL: HL in D. inornatus is 0.21, whereas it is 0.29 in D. hanagarthi . The temples in D. inornatus are 1.8 times as long as eyes, whereas they are only 1.3 times as long as eyes in D. hanagarthi . The seminal vesical is shorter than in D. inornatus and the coiled apical duct slightly larger.

Description: Length: 3.8 mm. Colouration: brown; pronotum yellow; elytra yellowish with indistinct brownish spot; antennae unicoloured dark yellow; legs yellow.

Head: 0.50 mm long, 0.43 mm wide; eyes slightly prominent; large; half as long as temples; EL: HL = 0.29; temples slightly divergent to posterior angles; PW: EW = 1.18; posterior angles widely rounded; not angulate; neck slightly shorter than half of head’s width; setiferous punctation deep; along wide impunctate midline with ca. 6 punctures in longitudinal line; temples laterally with moderately dense punctation; interstices between punctures approximately twice as wide as diameter of punctures; without microsculpture; surface shiny.

Antennae as long as head and half of pronotum combined; first antennomere nearly twice as long as second and slightly thicker; second and third antennomere conical and equally long; twice as long as apical width; following antennomeres distinctly shorter; approximately quadrate; basal 6 antennomeres with long apical setae; all antennomeres pubescent.

Pronotum: 0.62 mm long, 0.50 mm wide; sides nearly parallel; anterior and posterior angles widely rounded, not angulate; in dorsal aspect, fine lateral margin only visible shortly in front of posterior angles; wide midline impunctate; impunctate area much wider than lateral punctate area; few setiferous punctures in lateral area; setae long and thick; on average, interstices between punctures three to four times as wide as diameter of punctures; without microsculpture; surface shiny.

Elytra: 0.66 mm long, 0.66 mm wide; widest close to posterior angles; sides convergent to shoulders; shoulders distinct, but obtusely rounded; posterior margin triangularly emarginate; setiferous punctures in longitudinal lines with five to six punctures; on average, interstices between punctures four times as wide as diameter of punctures; setae long and thick, in particular, laterally; without microsculpture; surface shiny.

Abdomen densely pubescent.

Aedeagus with seminal vesical of endophallus two third as long as total length of central lobe; basal inner duct looped, apical inner duct with one large coil ending at apex; paramere reaching apex of central lobe; slender; nearly parallel from base to apex; only very slightly widened in apical part; without any process.

Spermatheca 0.42 mm long, 0.22 mm wide; bursa copulatrix in central position of approximately oval coiled duct; coiled duct widest at middle; sides in posterior half smoothly narrowed; anterior half nearly parallel.

Etymology: The species name honours the collector Dr. Werner Hanagarth, who unfortunately passed to early.

Figures

Figs 41: Fore-bodies of Diochus longicornis (A), D. tarsalis (B), D. adisi (C), D. ashei (D), D. guianensis (E), D. unicolor (F), D. inornatus (G), D. hanagarthi (H); scale bar: 0.5 mm.

Figs 6–9: Diochus longicornis-group: Diochus ashei (6), D. unicolor (7); Diochus inornatus-group: D. inornatus (8), D. hanagarthi (9); aedeagus (A), paramere (B) and antenna (C); scale bar: A, B: 0.1 mm, C: 0.2 mm.

Fig. 38: Spermathecae of Diochus longicornis-group: D. longicornis (A), D. tarsalis (B), D. ashei (C), D. guianensis (D), D. unicolor (E); Diochus inornatus-group: D. inornatus (F), D. hanagarthi (G), D. antennalis (lateral aspect H), D. hermani (I), D. hibbsi (J), D. newtoni (K), D. novus (L); scale bar: 0.1 mm.