Cenophengus brunneus Wittmer, 1976

Vega-Badillo, Viridiana, Morrone, Juan J. & Zaragoza-Caballero, Santiago, 2021, Revision of the genus Cenophengus LeConte, 1881 (Coleoptera, Phengodidae), with the description of four new species, new geographic records and a new synonymy, ZooKeys 1068, pp. 73-148 : 73

Cenophengus brunneus Wittmer, 1976

Fig. 5A-H

Cenophengus brunneus Wittmer, 1976: 453.

Type locality.

Veracruz, Mexico.

Type material examined.

Holotype ♂: "MEXICO: Veracruz/ Córdoba / Dr. A. Fenyes" " Cenophengus / Cenophengus brunneus det. W. Wittmer" “HOLOTYPUS” "Type No. / 73887/ USNM" | NMNH.

Remarks.

Cenophengus brunneus is sister to C. villae ( Vega-Badillo et al. 2021b), but can be distinguished by the interocular distance: in C. brunneus it is 3.5 times eye width, whereas in C. villae , it is 4.0 times longer. Additionally, in C. brunneus , the pronotal disc is convex, without a longitudinal carina, whereas in C. villae , it has a longitudinal carina.

Diagnosis.

Body brown, integument chagreened, antennae less than twice the length of the pronotum, antennal rami 1.5 times the respective antennomere and each elytron 5.1 times as long as wide; aedeagus with three teeth at the inner apex of paramere.

Redescription.

Male. Body length 4.30 mm; maximum body width 0.47 mm (pronotum). Body brown, legs paler (Fig. 5A, B ). Head. Wider (0.50) than long (0.40) (Fig. 5C ), at eye level, slightly wider than the pronotum, integument chagreened, punctures as large as eye facets and separated by approximately 1 punctured diameter, each puncture bearing a yellow-brown seta; interantennal distance (0.04 mm) less than length of the antennomere 1 (0.12 mm); eyes 1/2 as long as head in lateral view, longer (0.21 mm) than wide (0.09 mm); interocular distance (0.32 mm) 3.5 times eye width; short antennae (1.59 mm) less than twice the length of the pronotum; antennomere 1 (0.12 mm) as long as the next two combined, 3 cup-shaped, 4 (0.13 mm) shorter than the following antennomeres, 5 to 11 about equal in length (0.14 mm), 12 (terminal) (0.20 mm), antennal rami lanceolate in lateral view, 1.5 times the respective antennomere; terminal maxillary palpomere uniform, securiform (0.17 mm), as long as the preceding three combined (0.18 mm); terminal labial palpomere spindle-shaped (0.07 mm), 3 times as long as preceding one (0.02 mm). Thorax. Pronotum longer (0.64 mm) than wide (0.47 mm) (Fig. 5D ); integument chagreened, punctures as large as eye facets and separated by approximately 4 punctured diameters, each puncture bearing a yellow-brown seta; convex disc, weakly elevated dorsally forming a small depression in the basal part of each side, posterior margin curved with middle notch, sides almost straight, anterior angles rounded and posterior angles acute; mesosternal suture complete; scutellum with posterior margin rounded; each elytron 5.1 times as long (1.64 mm) as wide (0.32 mm), convex, without longitudinal costae, elytral apex rounded; hind wings with posterior radial vein (RP) length 3.8 times less than the length of MP1+2, radial cell closed, r3 vein present, r4 vein reduced (not reaching the RP or the radial cell), those of the anterior anal and posterior anal sectors, evident (Fig. 5E ). Legs: tarsomeres 1 and 2 of the prothoracic legs with a similar length and tarsomere 1 of meso- and metathoracic legs is longer than 2. Abdomen. Integument shiny, punctured, with long dense setae, sternite 7 with margin sinuate, sternite 8 with margin notched; aedeagus with three teeth at the inner apex of paramere (Fig. 5F-H ).

Female and immatures.

Unknown.

Distribution.

Mexico: Veracruz (Fig. 3 ).

Figures

Figure 3. Map of southern North America showing specimen localities for Cenophengus spp.

Figure 5. Cenophengus brunneus Wittmer, 1976, male. Habitus: A dorsal B ventral C head dorsal D pronotum dorsal E hind wing. Wing venation: CR = Radial Cell; r 3 = radial 3 vein; r 4 = radial 4 vein; RP = Posterior Radial vein; MP 1 + 2 = Posterior Median vein; CuA = Cubital vein; AA and AP = Anterior and Posterior Anal veins. Aedeagus: F dorsal view G lateral view H ventral view.