Lithocharodes peruana, Irmler, 2021

Irmler, Ulrich, 2021, The Neotropical species of the genus Lithocharodes SHARP, 1876 (Coleoptera: Staphylinidae: Staphylininae: Xantholinini), Beiträge Zur Entomologie = Contributions to Entomology 71 (1), pp. 29-85 : 54-55

Lithocharodes peruana spec. nov.

urn:lsid:zoobank.org:act:44E3E16F-6262-4A6B-8BF1-CE656372382A

Figs 7 a-d, 44 F, 48 C

Type material: male, holotype: Peru, Madre de Dios, Cocha Cashu Bio. Stn. , Manu National Park, (71°24'24"W, 11°53'45"S), 350 m elev., flight intercept trap, 17.- 19.10.2000, leg. R. Brooks, # PERU 1B00 042 ( KNHM) . Paratypes: 5 males, 2 females, with same data as holotype (6 KNHM, 1 UIC) ; Peru, Tambopata Prov. , Madre de Dios Dpt. , 15 km N Puerto, Maldonado Reserva Cuzco Amazónica (69°03'W, 12°33'S), 200 m elev., Plot #Z1U16, #Z1U19, #Z2U19, #Z2E14, #Z2E15, #Z2E16,Z1U16, #Z2U16, #Z2E17, #Z1 trail 26, Swamp trail, Quebrada Mariposa, flight intercept trap, 19 males, 40 females, 15.6., 17.6., 20.6., 22.6., 24.6., 26.6., 28.6., 3.7., 13.7., 15.7., 16.7., 17.7., 19.7., 30.6.1989, leg. J. Ashe, R.A. Leschen, #522 (53 KNHM, 6 UIC) ; same location and collectors but berlesate, Palmaceae flower fall/logs, Plot #Z2E20-21, male, 25.6.1999 ( KNHM) ; Cocha Salvador , Reserved Zone, Manu National Park (71°31'36"W, 12°0'13"S), 310 m elv., flight intercept trap, 5 males, 2 female, 20.- 21.10.2000, leg. R. Brooks, # PERU 1B00 070 (4 KNHM, 1 UIC) ; Cocha Cashu Biol. Stn. , Manu National Park (71°24'24"W, 11°53'45"S), 350 m elev., flight intercept trap, 1 male, 1 female, 17.- 19.10.2000, leg. R. Brooks, # PERU 1B00042 ( KNHM) ; Pantiacolla Lodge , 8 km N El Mirador trail, Alto Madre de Dios River (71°16'41"W, 12°38'30"S), 800 m elev., flight intercept trap, 23.- 26.10.2000, leg. R. Brooks, # PERU 1B00 102 ( KNHM) ; same location and collector but 5.5 km NW El Mirador trail, Alto Madre de Dios River (71°15'28"W, 12°39'10"S), 500 m elev., flight intercept trap, 3 females, 23.- 26.10.2000, # PERU 1B00100 ( KNHM) ; same region, CICRA Field Station , garden (70.10.10'W, 12.56.94'S), 260 m elev., malaise trap, 22.- 29.7.2010, leg. M. Endara, #PER10-07-MAT-008 ( KNHM) ; Dept. Loreto, Campamento San Jacinto (75°51.77'W, 2°18.75'S), 175-215 m elev., flight intercept trap, #66, female, 9.7.1993, leg. R. Leschen ( KNHM) .

Diagnosis: According to the small size, dark colouration and dense punctation of the head, L. peruana is similar to L. pampana and L. lescheni . The distinguishing characters are already listed there. Lithocharodes peruana can be recognized by the extremely divergent head. Males are easy to separate from other Lithocharodes species by the small, nearly circular aedeagus with a thick, transparent endophallus totally covered by minute teeth.

Description: Length: 4.3 mm. Colouration: Dark brown to blackish; legs and antennae lighter brown.

Head: 0.73 mm long, 0.63 mm wide; eyes large; slightly prominent; postocular space twice as long as eyes; postocular sides distinctly divergent; posterior angles widely rounded; posterior margin slightly convex; in short medial part straight; interantennal furrows long and deep; reaching posterior third of eyes; setiferous punctation deep and dense; partly coriaceous; on average, interstices between punctures less than half as wide as diameter of punctures; small midline impunctate; surface between punctures without microsculpture; shiny. Antennae with first antennomere as long as halflength of head; second and third antennomere conical and slightly longer than wide; subsequent antennomeres distinctly wider than long and increasing in width; fourth antennomere twice as wide as long; tenth antennomere 2.2 times as wide as long; all antennomeres densely pubescent. Pronotum: 0.85 mm long, 0.57 mm wide; widest in anterior third; anteriad, conically narrowed toward neck; slightly narrowed in middle third; approximately parallel in posterior third; posterior margin slightly convex; setiferous punctation as deep as on head but distinctly sparser; on average, interstices between punctures slightly shorter than diameter of punctures; moderately wide midline impunctate; midline in anterior third wider than in posterior third; surface without microsculpture; shiny. Elytra: 0.60 mm long, 0.55 mm wide; humeral angles sub-rectangular; sides divergent to posterior angles; posterior angles sub-rectangular; posterior margin slightly convex; retreated to suture; setiferous punctation weaker but denser than punctation on pronotum; on average, interstices between punctures half as wide as diameter of punctures; surface with weak isodiametric microsculpture; less shiny than pronotum. Abdomen with denser but weaker setiferous punctation than on elytra; isodiametric microsculpture deeper than on elytra; surface less shiny than elytra; posterior margin of sternite VII and tergite VII of male nearly straight. Aedeagus oval; endophallus thick, sacklike, covered by numerous minute teeth and folded several times; paramere bilobed; outer lobe simple without any setae; inner lobe moderately slender; slightly curved; apical part flexed; inner face with long setae; setae at base longer than setae at apex.

Etymology: The species name is derived from the coun-

try's name, where it was found.

Figures

Fig. 2–7: L.-verhaaghi-group: Lithocharodes verhaaghi (2), L. pampana (3), L. silvicola (4), L. trinitatis (5), L. dubiosa (6), L. peruana (7); aedeagus (a), paramere (b), sternite VII of male (c), tergite VII of male (d); scale bare: A – D: 0.2 mm; B: 0.1 mm.