Lithocharodes denticulata, Irmler, 2021

Irmler, Ulrich, 2021, The Neotropical species of the genus Lithocharodes SHARP, 1876 (Coleoptera: Staphylinidae: Staphylininae: Xantholinini), Beiträge Zur Entomologie = Contributions to Entomology 71 (1), pp. 29-85 : 40-41

Lithocharodes denticulata spec. nov.

urn:lsid:zoobank.org:act:9B194FB8-EFFE-4301-9681-174E24B19557

Figs 18 a-d, 44 M, 49 A

Type material: male, holotype: Bolivia, Cochabamba, Cochabamba, 67.5 km NE Est. Biol. Valle de Sajita, Univ. de San Simon (64°47'52"W, 17°6'33"S), 300 m elev., flight intercept trap, 9.- 13.2.1999, leg. F. Genier, #BOL1G99 069 ( KNHM) . Paratypes: Bolivia: 3 males, 3 females with same data as holotype ( KNHM) ; same location, date, and collecting method as holotype but 2 males, 2 females, leg. R. Hanley, #BOL1H99 078 (3 KNHM, 1 UIC) ; Peru: Madre de Dios, Cocha Salvador , Reserved Zone Manu National Park (71°31'36"W, 12°0'13"S), flight intercept trap, 4 males, 20.- 21.10.2000, leg. R. Brooks, # PERU 1B00 070 ( KNHM) ; Madre de Dios, Pentiacolla Lodge , Alto Madre de Dios River (71°13'55"W, 12°39'22"S), 400 m elev., flight intercept trap, 1 male, 1 female, 23.- 26.10.2000, leg. R. Brooks, # PERU 1B00 099 ( KNHM) ; same region, CICRA field station, ~ 2 km NW of cafeteria, research plot (70.1098°W, 12.5523°S), 295 m elev., flight intercept trap, pit fall trap, 5 males, 23 females, 7.-9.6., 9.-11.6., 11.-13.6., 12.- 13.6.2011, leg. Chaboo team, #PER-11-FIT-019 (25 KNHM, 3 UIC) ; Cusco Dept. , Villa Carmen field station, ~ 1.7 km west cafeteria research transect (71.4191°W, 12.8925°S), 555 m elev., flight intercept trap, 1 male, 3 females, 22.- 24.5.2001, leg. D.J. Bennett & E Razuri, #PER-11-FIT-009 ( KNHM) ; French Guiana: SaüL. 7 km N, Les Eaux Claires (53°13'19"W, 3°39'46"N), 220 m elev., flight intercept trap, male, 31.5.- 3.6.1997, leg. J. Ashe, R. Brooks ( KNHM) ; Mt Galbao summit (53°16'42"W, 3°37'18"N), 740 m elev., flight intercept trap, 2 females, 5.- 7.6.1997, leg. J. Ashe, R. Brooks ( KNHM) .

Diagnosis: The species is characterised by the apical row of large teeth of the endophallus. In addition, males can be identified by the specific shape of sternite VII with large lateral prominences and a wide medial prominence. Females are much more difficult to separate from similar species such as L. nigerrima . They are separated from similarly large species by the black colouration in combination with the sparse punctation of the posterior vertex, and the large eyes.

Description: Length: 4.4 mm. Colouration: Blackish; legs and antennae brown.

Head: 0.53 mm long, 0.52 mm wide; eyes large; slightly prominent; postocular sides slightly divergent; 1.8 times as long as eyes; interantennal furrows reaching anterior third of eyes; setiferous punctation moderately deep and dense; on average, interstices as wide as diameter of punctures; on anterior vertex interstices slightly shorter, on posterior vertex slightly wider; surface without microsculpture; shiny. Antennae with first antennomere slightly longer than half length of head; second and third antennomere conical; longer than wide; subsequent antennomeres wider than long and increasing in width; fourth antennomere 1.5 times as wide as long; tenth antennomere 2.2 times as wide as long; all antennomeres densely pubescent. Pronotum: 0.67 mm long, 0.44 mm wide; widest in anterior third; anteriad, conically narrowed toward neck; posteriad, slightly narrowed in middle third and nearly parallel in posterior third; posterior angles widely rounded; posterior margin convex; setiferous punctation as dense and deep as on posterior vertex; wide midline impunctate; surface without microsculpture; shiny. Elytra: 0.62 mm long, 0.56 mm wide; humeral angles obtuse; sides posteriad nearly parallel; posterior angles sub-rectangular; posterior margin slightly convex with wide incision at suture; setiferous punctation finer than on pronotum but slightly denser; surface with extremely fine isodiametric microsculpture; shiny. Abdomen with denser and finer setiferous punctation than on fore-body; microsculpture denser and deeper than on elytra; surface less shiny; sternite VII of male with long curved lateral teeth; posterior margin deeply emarginate on each side of middle; central prominence convex; tergite VII of male with straight posterior margin. Meso- and metatibia with three subapical ctenidia. Aedeagus long-oval with sub-rectangular anterior angles; posterior part of endophallus covered by minute teeth; in anterior part with row of conspicuously large teeth; paramere short and broad; bilobed; outer lobe broad; triangular and half as long as inner lobe; inner lobe with straight inner face and slightly curved outer face; sparsely covered by fine setae; setae on inner face denser; basal setae slightly thicker and longer than apical setae.

Etymology: The species name is derived from the same Latin word (meaning toothed) and refers to the shape of the male sternite VII.

Figures

Fig. 13–18: L.-simillima-group:Lithocharodes triangula (13), L. simillima(14), L. surinamensis (15), L. brooksi(16), L. germana (17), L. denticulata (18); aedeagus (a), paramere (b), sternite VII of male (c), tergite VII of male (d); scale bare: A–D: 0.2 mm; B: 0.1 mm.