Coprophanaeus, Olsoufieff, 1924

Edmonds, W. D. & Zidek, J., 2010, A taxonomic review of the neotropical genus Coprophanaeus Olsoufieff, 1924 (Coleoptera: Scarabaeidae, Scarabaeinae), Insecta Mundi 2010 (129), pp. 1-111 : 48-50

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Coprophanaeus (C.) jasius ( Olivier, 1789)

Fig. 96 View Figure 95-101 , 103, 107, 109 View Figure 102-112 , 113 View Figure 113 , 129-133 View Figure 129-133

Scarabaeus jasius Olivier, 1789: 109

Phanaeus satyrus Castelnau, 1840: 80 (syn. by Nevinson 1891: 1 as P. acrisius )

Coprophanaeus jasius (Olivier) (recomb. by Blackwelder 1944: 209)

Type. S. jasius – neotype male (des. by Arnaud 2002a: 7), Muséum National d’Histoire Naturelle, Paris (examined by photo); P. satyrus – unknown to us.

Diagnosis. General – Metallic coloration green to yellowish green (seldom with red reflections) confined to posterior areas of head and pronotal sides and margins. Clypeal margin ( Fig. 129, 132 View Figure 129-133 ) clearly explanate/ angulate lateral to median teeth. Posteromedian portion of pronotum ( Fig. 96 View Figure 95-101 ) smooth, bearing sparse, effaced puncturing (weakest in large males). Sulcus paralleling carinate posterior margin of pronotum smooth ( Fig. 109 View Figure 102-112 ), lacking conspicuous puncturing or asperities (x25); carina entire. Posterior pronotal fossae with sparse puncturing and/or microgranules; area surrounding fossae virtually smooth. Length 18-27 mm.

Male ( Fig. 96 View Figure 95-101 , 103 View Figure 102-112 , 131-133 View Figure 129-133 ) – Tip of head horn never reaching level of prothoracic carina; anterior surface of base usually coarsely punctate; central process (viewed laterally) inclined slightly anteriorly from base, its length less than one-half that of base. Pronotal carina ( Fig. 96 View Figure 95-101 ) weakly, broadly angulate medially, trituberculate, medial tooth only slightly larger than laterals. Parameres long, length at least three-fourths that of phallobase.

Female ( Fig. 107 View Figure 102-112 , 129-130 View Figure 129-133 ) – Cephalic carina bowed or angulate medially, median tooth larger than laterals. Vertex coarsely, densely punctured. Pronotal ridge trapezoidal, summit weakly trituberculate, width at summit slightly less than interocular width. Anteromedian concavity shallow, posterior (upper) margin simple, not bitumose.

Specimens examined – 274.

Distribution. Amazonian subregion ( Fig. 113 View Figure 113 ).

Collection Records. BRAZIL: Amazonas – 26 km N Manaus [Reserva Forestal Ducke] (Oct-Dec) ; 60 km N. Manaus, Fazenda Esteio and Fazenda Dimona (Jan, Jun-Jul); Municipio Itacoatiara, Fazenda Aruana [AM Hwy 10, km 215] (Nov); Manacapurú (Mar). Pará – Belêm [Agua Preta] (Jan-Feb]; upper Rocana River. Rondônia – 9 km NE Cacaulândia (Feb) . FRENCH GUIANA: 7 km N Saul, 3 km NW Les Eaux Claires, Mt. La Fumée , 490 m, 3 o 39’46”N 53 o 13’19”W (Jun) GoogleMaps ; Saül, Mt. Galbao , 3 o 37’18”N 53 o 16’42”W 740 m (Jun) GoogleMaps ; Nouragues Field Station , 4 o 05’N 52 o 40’W GoogleMaps . GUYANA: Cuyuni-Mazaruni – Bartica District (May). Potaro- Siparuni – Iwokrama Forest Reserve, 4 o 40’19”N 58 o 41’04”W, 100-200 m (May-June) GoogleMaps . SURINAME: Sipaliwini – Central Suriname Nature Reserve, environs of Voltzberg research station, 4 o 40.90’N 56 o 11.13'’W, 100 m (Mar) ; Central Suriname Nature Reserve, environs of Lolopasi field station, 4 o 42.91’N 56 o 12.83'’W, 80 m (Mar) ; Central Suriname Nature Reserve, environs of Conservation International field station approx. 50 m (Mar) ; Oelemarie, ~ 3 o 6’0'’N 54 o 32’0'’W (Aug); Palumeu, ~3 o 21’30.3'’N 55 o 26’38.2'’W (Jun). VENEZUELA: Bolívar – Lago Guri Islands , 7 o 21”S 62 o 52’W (Jun) ; Las Trincheras [ Río Caura ] (Aug) ; Puerto Cabello [ Río Caura ] (Aug) ; Jabillal [ Río Caura – Orinoco] (Oct). Delta Amacuro – 11 km W Piacoa.

Comments. There has been much confusion attending the use of the name “ jasius ”; it has been regularly and erroneously applied to practically all members of the jasius group. Arnaud’s (2002a) designation of a neotype resolved much of the identity problem of this, the type species of the genus. Coprophanaeus jasius is an Amazonian species commonly found in the same communities as its congeners C. parvulus , C. dardanus and C. lancifer . It is a copronecrophagous forest dweller. Its closest known relative is C. abas , originally described by MacLeay as a variety of C. jasius and first recognized as a separate species by Arnaud (2002c). Of the members of the jasius group, only C. jasius and C. abas have a conspicuously smooth posteromedian portion of the pronotum.

Pereira and Martínez (1956; see also 1960) synonymized C. cyanescens and C. camargoi with C. jasius , but neither is now considered in the context of Olivier’s taxon. We consider C. cyanescens (q.v.) a valid species and C. camargoi a synonym of C. acrisius (q.v.). Arnaud (2002c) has a wider definition of C. jasius than our own; he cites the species from Colombia, Ecuador, Peru and Bolivia. While we have seen no specimens from any of these countries, its occurrence in Amazonian habitats in any or all of them is not out of the question and we so indicate in the distribution depicted in Fig. 113 View Figure 113 .

Arnaud (1982) pointed out that the type of Phanaeus costatus is in fact a “hybrid” combining the forebody of a Coprophanaeus with the hindbody of Diabroctis mimas (Linné) . He identified the forebody as C. jasius , which he synonymized with C. costatus . C.P.D.T. Gillett (pers. comm.) has kindly informed us that the forebody is of C. cyanescens , requiring that the synonymy be reassigned accordingly (see Comments under C. cyanescens ).













Edmonds, W. D. & Zidek, J. 2010

Coprophanaeus jasius (Olivier)

Blackwelder, R. E. 1944: 209

Phanaeus satyrus

Nevinson, B. G. 1891: 1
Castelnau, F. L. 1840: 80

Scarabaeus jasius

Olivier, A. 1789: 109