Coprophanaeus, Olsoufieff, 1924

Edmonds, W. D. & Zidek, J., 2010, A taxonomic review of the neotropical genus Coprophanaeus Olsoufieff, 1924 (Coleoptera: Scarabaeidae, Scarabaeinae), Insecta Mundi 2010 (129), pp. 1-111 : 38

Subgenus Coprophanaeus s. str. Olsoufieff, 1924

Coprophanaeus Olsoufieff, 1924: 22 . Type Species: Scarabaeus jasius Olivier , original designation.

Diagnosis. General – Lower portion of eye large, width greater than twice (usually about three times or more) that of oculogular space. Paraocular areas (genae) not carinate lateral to eyes. Occipital areas of parietals more-or-less flattened, lacking angulate prominence ( Fig. 12 ). Circumnotal ridge entire or broken behind eyes ( Fig. 7-8 ). Pronotal sculpturing variable, usually granulorugose anterolaterally, punctate to some degree posteromedially; basal fossae variable. Pronotum lacking posterolateral ridge. Striae fine, simple or carinulate. Posterior edge of hind wing strongly notched basally ( Fig. 9 ). Posterior surface of each protibial tooth with single basal row of long setae ( Fig. 6 ). Abdominal sterna usually smooth medially. Sexual dimorphism marked.

Male – Head bearing lamellate process of some sort or trituberculate carina, never a simple curved horn. Pronotum variable, flattened or transversely concave anteromedially, with strong median prominence. Parameres variable.

Female – Head with trituberculate carina, sometimes strongly raised. Protarsi absent. Pronotum variable, usually with transverse process near anterior margin followed by oval concavity.

Distribution. Neotropical region.

Comments. This subgenus embraces 29 species distributed in four species groups separated in the key below. Edmonds (1972) divided the subgenus into two species groups (jasius and dardanus); only the jasius group is maintained here as defined then. Our organization of this subgenus is different from that presented by Arnaud (2002c), but many of the subgroups of closely related species (Arnaud’s complexes) are the same. Keys to species appear under the treatments of each species group.

Figures

Figure 5-12. Characters of Coprophanaeus. 5) Ventral view of front left tibia of C. ensifer (arrow marks elongate tuft of setae). 6) Same, C. dardanus. 7) Left anterolateral pronotal margin of C. spitzi (arrow indicates unbroken circumnotal carina. 8) Same, C. magnoi (arrow indicates broken circumnotal carina). 9) Base of left hind wing of C. cerberus. 10) Same, C. horus. 11) Lateral view left occipital lobe of C. ensifer (asterisk marks swelling). 12) Same, C. spitzi (asterisk marks flat area).