Manota sigma, Kurina, Olavi & Hippa, Heikki, 2015

Kurina, Olavi & Hippa, Heikki, 2015, A review of the South Pacific Manota Williston (Diptera, Mycetophilidae), with the description of thirteen new species, Zootaxa 4020 (2), pp. 257-288 : 280-282

Manota sigma sp. n.

Figs 13 A–F

Male. Colour. Head brown, face somewhat paler. Antenna including scape and pedicel brown, two apical flagellomeres somewhat paler. Clypeus and mouthparts pale yellowish. Thorax light brown, medial part of scutum and scutellum darker. Legs yellowish, basal fourth of hind coxa light brown, basal half of mid and hind femora infuscated. Wing hyaline; haltere yellow with dark brown knob. Abdomen light brown, medial 2/3 of tergites darker. All setosity pale, yellowish or brownish, the thicker setae seeming darker than the finer ones. Head. Antennal flagellomere 4 Fig. 13 A. Palpal segment 3 of maxillary palpus with apicomedial thumb-like extension, with 3 apically curved and somewhat flattened sensilla; palpal segment 4 subapically constricted, without parasegment; palpal segment 5 ~1.8 times longer than palpal segment 4. Nine strong postocular setae. Thorax. Anepisternum with 43–45 setae; anterior basalare non-setose; preepisternum with 9 setae; laterotergite with 29 setae; metepisternum with 12 setae. Legs. Mid- and hind tibial organs absent. Wing. R1 meeting C within the basal half of the costal margin; the sclerotized part of M2 proximally faint, hardly extending to the level of the tip of R1; wing length 2.2 mm. Hypopygium, Figs. 13 B–F. Sternite 9 small, subquadrangular, less than half the ventral length of gonocoxa, laterally separated from gonocoxa, anteriorly deeply incised, posteriorly slightly notched, the setae uniform and similar to the ventral setae of gonocoxa. Ventromedial margin of gonocoxa slightly sigmoid. Parastylar lobe weakly sclerotized, appearing as a small plate at the medial gonocoxal margin, with two long setae. Paraapodemal lobe small, only partly visible in the ventral view. Dorsomedial margin of gonocoxa slightly concave, with a small setose thumb-like lobe on posterior half. Posteriorly at the dorsal medial margin of the gonocoxa, on more ventral level with a subquadrangular oblique lobe bearing numerous setae at its apical margin and ventral surface. Dorsal gonocoxal setae unmodified. Gonostylus elongate, as long as gonocoxa, slightly sigmoid, setae uniform except these at the medial margin elongated, apical part of ventral surface and basal part of dorsal surface widely non-setose. One juxtagonostylar seta present as an apically flattened and expanded megaseta arising from a low basal body that is not more than one third of the length of the megaseta. Aedeagus subtriangular, with well expressed lateral shoulders, the apex curved ventrad. Hypoproct posteriorly extending to the basal part of gonostylus, its ventral part (sternite 10) with ~23 setae on each half, the setae distributed on medial and anterior marginal areas. Cerci medially separate, setose apically and laterally.

Female unknown.

Discussion. Manota sigma is similar to M. kaindiensis and runs also to couplet 14 in Hippa (2007) but see specific discussion under the latter for characters unique to M. sigma .

Etymology. The name sigma is derived from the eighteenth letter of the Greek alphabet and refers to the sigma-shaped gonostylus.

Types. Holotype. Male, PAPUA NEW GUINEA, SE, Koroba, 40 km W of Tari, 1650 m, light trap, 18.ix.1963, R. Straatman (BPBM).

Figures

FIGURE 13. Manota sigma sp. n. (holotype). A. Antennal flagellomere 4, lateral view. B. Hypopygium, dorsal view. C. Hypopygium, ventral view. D. Gonostylus, dorsal view. E. Cerci and apex of aedeagus, dorsal view. F. Hypopygium, aedeagus and posteromedial lobe on the dorsal side of gonocoxa, ventral view. Scale 0.1 mm.