Trichophallus hamatus, Ingrisch, 2024

Ingrisch, Sigfrid, 2024, Revision of the genus Trichophallus Ingrisch, 1998 with notes on the genera Secsiva Walker, 1869 and Subrioides C. Willemse, 1966 (Orthoptera: Tettigoniidae: Conocephalinae: Agraeciini), Zootaxa 5442 (1), pp. 1-66 : 26-28

Trichophallus hamatus sp. nov.

Fig. 10

Subria gracilis [nec Karny 1907] Karny 1912, Abh. Ber. Zool. Mus. Dresden 14(2): 7 (misidentification).

Holotype (male): Papua New Guinea: Toricelli mountains , 600 m (3°30‘S, 142°0‘E), I.1910, Dr. Schlaginhaufen (Dresden, SMTD).

Other specimens studied: Papua New Guinea: Toricelli mountains , 120–900 m (3°30‘S, 142°0‘E), I.1910, Dr. Schlaginhaufen — 1 male, 5 females , paratypes (Dresden, SMTD); West Sepik, Feramin , elev. 1450 m (5°12‘15.16‘‘S, 141°44‘11.89‘‘E), 30– 31.viii.1963, coll. R. Straatman — 1 male (Honolulu, BPBM); Telefomin, elev. 1600 m (5°10‘S, 141°35‘E), 30.viii.1963, coll. R. Straatman — 1 male (Honolulu, BPBM) .

Remark. The specimens had been originally labeled as „ Subria gracilis det. Karny“

Diagnosis. T. hamatus is characterised by the shape of the male cerci that are elongate, narrow, and regularly curved from base to obtuse tip and carry in subapical area from internal margin a short, acute spine. The titillators have the apical areas moveable against the basal area; the apical areas each carry a lateral membranous bursa with setose margin and they divide at end into a larger sickle-shaped and a shorter obtuse lobe. The female subgenital plate is more strongly modified than in other species of the genus with the central plate reduced to a tiny cone and the basal-lateral extensions are strongly prolonged into a pair of somewhat wavy and partly widened lobes.

Description. Medium sized. Fastigium verticis conical, shorter than scapus, apex obtuse; ventral margin compressed, separated by a shallow excision from fastigium frontis. Frons shining, subsmooth, with few, very shallowly impressed dots. Pronotum with lateral angles rounded, only apical area shouldered and disc flat; surface shining with few impressed dots throughout but auditory swelling and flat apical area of disc often subrugose and with a low step at begin of flat apical area; anterior margin broadly rounded but subtruncate in middle; posterior margin rounded; first transverse sulcus little distinct, second sulcus distinct only on paranota. Paranota much longer than high; ventral margin almost straight with ventro-posterior angle little produced. Fully winged; tegmen narrow, reaching about apical third of hind tibia. Prosternum with two minute tubercles, without spines. Meso- and metasternal lobes rounded; medial plate with a spine at both posterior angles. Mesocoxa with a small spinule. Femora with the following number of spines on ventral margins: fore femur 6–9 external, 5–7 internal; mid femur 7 external, 3–4 internal near base; hind femur 10–12 external, 9–15 internal over the whole length. Knee lobes of fore and of mid femur obtuse on external, obtuse or triangular on internal side; of hind femur bi-spinose on both sides. hind tibia with 1 dorsal and 2 ventral apical spurs on each side. Anterior tibia with dorsal angles rounded.

Male. Stridulatory file with about 114–135 teeth. Tenth abdominal tergite transverse, in midline slightly sunk in; apical margin on both sides of middle a little swollen and angularly extended, terminating into two very short, obtuse, projections, truncate or slightly concave in between. Epiproct rounded but apical area triangular with apex obtuse; with a pit or short furrow in middle of basal area. Paraprocts with a rather long, slightly curved, obtuse projection from internal margin. Cerci long and narrow, slightly curved, conical, with an obtuse, pre-apical, internal tooth; apex obtuse. Subgenital plate with apical margin roundly excised.

Titillators compressed, small; brown in lateral and apical, hyaline in central-basal areas; base and central area broad, in apical third suddenly constricted and apex conical; at the point where the titillators are constricted, a pair of separate, elongate sclerites is inserting which support each a hyaline fold of the phallus and divide at end into a larger sickle-shaped and a shorter obtuse process. The lateral hyaline folds are roughly disc-shaped, hollow, and carry at the apical-internal margin a row of stout bristles and at the opposite margin a field of short, fair hairs.

Female. Terminal tergite transverse, split in midline; apical margin substraight but excised in middle. Cerci long conical, slender, slightly curved, apex pointing. Subgenital plate divided into two compressed, sinusoidal curved lateral lobes with obtuse apex, only at very base connected by a small and very short central lobe which is roundly produced behind and has a medial carina; ventral surface of lateral lobes setose. Ovipositor elongatefalcate, compressed in apical half, apex acute; dorsal margin of ventral valves swollen at base and ventral surface of this swollen area setose.

Coloration. Uniformly yellowish brown. Tegmen with some scattered faint brown dots especially in anterior area before subcosta.

Measurements (4 males, 5 females).—body w/o wings: male 20–23, female 22–23; pronotum: male 5.3–6.2, female 5.8–6.2; tegmen: male 31–33, female 33.5–36; hind femur: male 15.5–16.5, female 16.5–18.5; ovipositor: female 10.5–11.5 mm.

Etymology. The name of the new species refers to the nearly hook-shaped apex of the male cerci; from Latin hamatus = hooked.

Figures

FIGURE 10. Trichophallus hamatus sp. nov.: A, male holotype; B, stridulatory file; C–F, titillators in different aspects (C–E, male from Feramin; F, male from Toricelli Mts); G–H, last abdominal tergite and cerci in dorsal view (G, of holotype); I, tip of male cercus with apical spine; J, male epiproct and paraprocts in dorso-apical view; K, male subgenital plate and cerci in ventral view; L, face; M, female habitus in ventral view; N, female subgenital plate of allotype. Scales: A 10 mm, B–E 1 mm, H and K 5 mm.