Trichophallus furcatus, Ingrisch, 2024

Ingrisch, Sigfrid, 2024, Revision of the genus Trichophallus Ingrisch, 1998 with notes on the genera Secsiva Walker, 1869 and Subrioides C. Willemse, 1966 (Orthoptera: Tettigoniidae: Conocephalinae: Agraeciini), Zootaxa 5442 (1), pp. 1-66 : 21-23

Trichophallus furcatus sp. nov.

Fig. 7

Holotype (male i33): Indonesia, Maluku, Kei Islands, Gunung Daab (5°35‘10‘‘S 133°4‘4‘‘E), 300m, IV.1922, H.C. Siebers ( MZB, Bogor Java).

Other specimens studied: Paratypes: Indonesia, Kei Isl. 1922, Gunung Daab, H.C. Siebers ( MZB, Bogor Java) : 2 males, no. 94; 1 female, no. 100; 4 females, no. 100, 112, 134, and 138.

Diagnosis. The new species is characterised by the shape of the male cerci with substraight trunk, incurved apical area with approaching margins and obtuse tip, and a narrow, styliform projection arising from the curvature. The shape of the titillators with four sclerites is also characteristic. The shape of the elongate female subgenital plate that has the ventral area forming a pair of rather wide, convex and long ventral apical lobes with approaching margins is also characteristic for this species.

Description. Fastigium verticis laterally compressed; apex obtusely rounded. Frons shining with a few shallow, hardly impressed dots. Pronotum semi-cylindrical, apical area of disc nearly flat and shouldered; transverse sulcus weak, interrupted in middle; a second sulcus behind middle of pronotum is only distinct on paranota; anterior margin broadly rounded but slightly concave in middle; posterior margin very faintly rounded, almost subtruncate, sometimes faintly incised in middle; ventral margin descending posteriorly, rounded angular below apical callus, ascending afterwards; humeral sinus hardly indicated. Tegmen long and narrow, largely surpassing apex of ovipositor and hind knees. Prosternum with 2 very small tubercles, more distinct in female than in male. Meso- and metasternal lobes rounded or angularly rounded; medial plate with a strong spine at each apical angle. Mesocoxa with ventral internal projection rather long (less so on postcoxa), and with a distinct spinule on dorsal-anterior margin and sometimes with a 2nd spinule somewhat below. Femora with the following number of spines on ventral margins: fore femur 7–9 external, 5–7 internal; mid femur 7–8 external, 2–4 internal near base; hind femur 11–16 external, 12–18 internal (external spines larger than internal spines). Knee lobes of fore femur obtuse to triangular on external, spinose on internal side; of mid femur obtuse-triangular on external, triangular on internal side, of hind femur bi-spinose on both sides (the second smaller spine sometimes lacking, sometimes also present on medial femur). Hind tibia with 1 + 1 dorsal and 2 + 2 ventral, external and internal, apical spurs.

Male. Stridulatory file with about 107 teeth. Tenth tergite slightly prolonged behind; apical margin concave between 2 lateral short-conical, obtuse projections. Epiproct rounded triangular, dorsal surface shallowly depressed, lateral margins sloping in a more than 90°-angle. Cerci semi-cylindrical on external side, depressed on internal side; depressed area terminating into a large subapical internal tooth with acute apex; external area terminating in a styliform projection with obtuse apex. Subgenital plate broad, lateral margins sloping in basal area; with or without a faint medial carinula before apex and with a strong rounded carina on each side before base of styli; apical margin roundly emarginate between bases of styli.

Titillators simple but covered by complex sheaths in form of 2 pairs of membranous bursae. Apex of titillators connected via a less sclerotised area with a subflat sclerite which supports the internal area of the dorsal bursa; this sclerite with a lateral hook that is fused with the surrounding membrane except for the apex which is free. The ventral bursae paired; compressed-ovoid in general outline with the lateral margin densely covered with bristles. Dorsal bursae compressed and slightly curved; surface granulose, slightly wrinkled and apex covered densely with short hairs. The whole apex of the titillator complex is embraced by another semi-cylindrical membranous sheath, which is granulose, wrinkled, and double-fold.

Female. Tenth tergite furrowed in midline, apical margin subtruncate on both sides, angularly excised in middle. Epiproct rounded-triangular, disc depressed. Cerci long-cylindrical, gently curved, apex pointing. Subgenital plate elongate, about apical half split into two long-conical compressed lobes with obtuse apices; the entire basal area with a club-shaped groove in middle and a lateral carina at each side; with basal-lateral sclerites that have a bulging apical-lateral margin and are furrowed in middle. Ovipositor falcate; with margins hardly narrowing before apical third, of sub-equal height in basal two thirds, narrowing in apical third; margins smooth; ventral valves faintly longer than dorsal valves.

Coloration. Yellowish-brown. Fastigium verticis, vertex and disc of pronotum with a narrow, indistinctly brown, medial band, interrupted in midline by a pale line. Frons in some specimens slightly darkened around medial ocellus. Tegmina with dark brown dots. Legs with a few indistinct spots. Spines partly darkened.

Measurements: body male 26–28, female 26–28; pronotum male 6.5–7.2, female 6.7–7.2; tegmen male 33–36, female 37–40; hind femur male 17–19, female 19–21; ovipositor 12.0– 12.5 mm.

Etymology. The name of the new species is deviated from the Latin word furcatus forked, refering to the shape of the male cerci.

Figures

FIGURE 7. Trichophallus furcatus sp. nov. male (A–H) and female (I–J): A, habitus lateral view; B, face; C, subgenital plate and cerci ventral view; D–E, last tergite and cerci in obliqe dorsal (D) and strictly dorsal (E) view; F–H, phallus with titillators and membranous structures in lateral view (F), in ventro-apical view with membranous structures spread apart (G) and in situ with membranes embracing titillators (H); abdominal apex with ovipositor and subgenital plate in lateral view (I); subgenital plate in ventral view (J). Classical photography, not to scale.