Trichophallus forcipatus, Ingrisch, 2024

Ingrisch, Sigfrid, 2024, Revision of the genus Trichophallus Ingrisch, 1998 with notes on the genera Secsiva Walker, 1869 and Subrioides C. Willemse, 1966 (Orthoptera: Tettigoniidae: Conocephalinae: Agraeciini), Zootaxa 5442 (1), pp. 1-66 : 18-20

Trichophallus forcipatus sp. nov.

Figs. 5–6

Holotype (male): North New Guinea, Mamberamo , Albatros Bivak, (2°17‘S, 138°1‘E), 1–31.v.1926, coll. W. Docters v. Leeuwen, North New Guinea Exped. ( MBBJ Bogor).

Other specimens studied: Indonesia: Papua, Jayawijaya (Central) Mountains, Archbold Lake, elev. 760 m (3°41‘S, 138°53‘E), 26.xi.–3.xii.1961, coll. S. & L. Quate — 1 male ( BPBM Honolulu); Mamberamo Area , Pionierbivak, (2°20‘S, 138°0‘E), 1–30.vi.1926, coll. W. Docters v. Leeuwen, North New Guinea Exped.— 1 female ( MBBJ Bogor) .

Diagnosis. Male cerci with dorsal margin convex, curved throughout, ventral margin with swollen external, flattened internal margin; from internal margin with a little curved, triangular projection, and with a second subsinuate projection from internal apical margin. Titillators with apical areas, carrying hairs, clearly separated from fused basal-central areas.

Remark. There are minor differences in the male characters between specimens from different localities. Thus it might be possible that the species could be divided into two subspecies when more specimens become available for study.

Description. Rather large for the genus. Fastigium verticis laterally compressed, dorsal surface very faintly furrowed, apex obtusely rounded. Frons shining with hardly any impressed dots. Pronotum semi-cylindrical, apical area of disc almost flat and shouldered; transverse sulcus weak, interrupted in midline; anterior margin broadly rounded but slightly concave in middle; posterior margin faintly rounded, almost subtruncate; ventral margin descending backwards, rounded below apical callus and afterwards ascending; humeral sinus faintly indicated. Tegmen long and narrow, surpassing middle of stretched hind tibia. Prosternum with 2 very faint, hardly visible tubercles. Meso- and metasternal lobes rounded; medial plate with an obtuse conus at each posterior angle. Mesocoxa with ventral internal projection rather long (less so on postcoxa), dorsal-anterior margin with a weak, obtuse-angular projection. Femora with the following number of spines on ventral margins: fore femur 7–8 external, 5–7 internal; mid femur 7–8 external, 3–4 internal near base; hind femur 12–15 external, 12–16 internal (the basal spines very tiny). Knee lobes of fore femur obtuse on external, spinose on internal side; of mid femur obtuse on external, triangular but apex obtuse on internal side; of hind femur bi-spinose on both sides. hind tibia with 1 dorsal and 2 ventral apical spurs on each side.

Male. Stridulatory file with circa 122–136 teeth. Tenth tergite prolonged behind; hind margin roundly incised in middle between two short obtuse-angular lobes; setose in central and apical areas. Epiproct angularly rounded, with a Y-shaped carina between disc and sloping lateral area. Paraprocts with apical projections rather long, finger-shaped, curved dorsad. Cerci slightly curved mediad; dorsal external area rounded, ventral internal area flattened or little concave; with a large, depressed, about triangular internal projection in circa middle of cercus length terminating into an obtuse conus, and also with an elongate-conical, S-shaped ventral apical process; very apex of main cercus trunk rounded. Subgenital plate with sloping lateral area rather low; lateral margins carinated, disc with faintly rounded carinae before bases of styli; apex broad-roundly emarginated; styli rather long and stout.

Titillators simple, apical parts fused, but very apices separate; connected with complex sheaths. Each apical part connected via a less sclerotised area with another apically rounded plate supporting the apical sheath on internal side. Sheaths bursa shaped, in dorsal-apical view ring-shaped embracing apices of titillators; somewhat darkened and granulose on external surface; internal surface hyaline; on ventral internal surface in apico-lateral areas with a curved row of long bristles, the longest of them surpassing the apex of the sheath. Laterally of the sheath the surrounding membranes forming another hyaline bursa. Membrane between bases of titillators with a button-shaped protuberance.

Female. Subgenital plate elongate with converging lateral margins, divided from hind margin along mid-length into two lobes that are curved ventrad at end and reach about the elongate basal lobes of the ventral ovipositor valves.

Remark. The female has been tentatively assigned to this species since it has not been collected with either of the two males, although at the same river system as the holotype.

Coloration. Uniformly yellowish-brown. Tegmen with few hardly darkened dots. Tibiae with a small, subbasal, dark spot; fore tibia with 2 spots: above and below tympanum.

Measurements (2 males): body: male 23–28; pronotum: male 5.8–6.2; tegmen: male 36–38; hind femur: male 18.5–19.7; antenna: male 95 mm. Measurements of female missing.

Etymology. The name of the new species is deviated from Latin forcipata = scissors.

Figures

FIGURE 5. Trichophallus forcipatus sp. nov. males from Archbold Lake (A–G) and from Mamberamo (H–K).—A, stridulatory file; B, subgenital plate and Cerci ventral view; C–F, titillators in apical (C), ventro-distal (D), dorso-proximal view (E), view on tip (F); G–H, tenth tergite and cerci; I–K, titillators in ventro-distal (I), dorso-proximal (J), and apical (K) view.—Scales for titillators about 1 mm.

FIGURE 6. Trichophallus forcipatus sp. nov. female from Pionierbivak.—A–C, subgenital plate in ventral (A), in apical (B) and in lateral view (C); D, end of abdomen and base of ovipositor with basal lateral lobe in lateral view; E, ovipositor; F, subgenital plate and base of ovipositor in ventral view.