Diplommatina auriculata Möllendorff, 1897

Hausdorf, Bernhard, 2017, Revision of the Diplommatinidae (Gastropoda: Cyclophoroidea) from Java, Zootaxa 4312 (2), pp. 201-245 : 228-235

Diplommatina auriculata Möllendorff, 1897

( Figs 2 , 5 , 17 , 41–44 ; Table 1)

Diplommatina (Sinica) auriculata Möllendorff, 1897: 95 (JaVa) . Kobelt 1902: 456. LeSchke 1914: 215. RenSch, 1931: 386, fig. 11. RenSch 1934: 744. ParaVicini 1935: 174.

Diplommatina (Sinica) auriculata Var. abbreviata Möllendorff, 1897: 96 (JaVa) (not Heude 1885). Kobelt 1902: 456.

Diplommatina (Sinica) auriculata Var. biangulata Möllendorff, 1897: 96 (JaVa). Kobelt 1902: 456.

Diplommatina (Sinica) auriculata Var. gracilior Möllendorff, 1897: 96 (JaVa). Kobelt 1902: 456.

Diplommatina (Sinica) hortulana — LeSchke 1914: 215, 285 (in part, not LeSchke 1914).

Diplommatina auriculata — Van Benthem Jutting 1932: 202, fig. 5–7 (See Van Benthem Jutting 1940). Van Benthem Jutting 1940: 332; 1948: 593, fig. 46, 50. Vermeulen & Whitten 1998: 47, 137, fig. 24. Heryanto et al. 2003: 85, fig. Heryanto 2008: 362 (partly). Heryanto 2009: 332 (partly). Heryanto 2011: 45, figS. Heryanto 2012: 196.

Diplommatina (Sinica) auriculata auriculata — Zilch 1953: 25, pl. 12 fig. 173–176.

Diplommatina cyclostoma —Heryanto 2008: 362 (partly, not Möllendorff 1897). Heryanto 2009: 332 (partly, not Möllendorff 1897).

Diagnosis. Diplommatina auriculata is characterized by a medium-sized shell with distinct extensions of the peristome, a prominent columellar lamella and close-set ribs.

Shell ( Fig. 2 , 5 , 41–44 ; Table 1). Dextral; spindle shaped with conical early whorls and a narrowed last whorl; with 6.5–8 whorls separated by a deep suture; protoconch smooth; teleoconch with oblique ribs (28–92 on the penultimate whorl); with or without discernible spiral striae between the ribs; corneous; subtranslucent; last whorl rounded, with a ventral constriction; aperture subangular, angular at the transition from columellar to basal side; upper insertion of the peristome distinctly ascending towards aperture; insertions of the peristome connected by a distinct callus; peristome distinctly expanded, partly with 1–4 blunt processes, thickened, doubled; aperture with a prominent columellar lamella; with a parietal lamella ventrolaterally left inside the last whorl, sometimes with two parietal lamellae in one line, separated by a gap; with a short spiral palatal fold in the middle of the whorl ventrolaterally left inside the last whorl, sometimes with an additional very short spiral fold close to the suture; with a radial palatal fold ventrally; umbilicus closed; operculum thin, corneous, circular, paucispiral. Shell height 2.9–4.45 mm, shell diameter 1.45–2.2 mm.

Remarks. Diplommatina auriculata differs from D. canaliculata in the less slender spire and the close-set ribs (28–92 on the penultimate whorl versus 25–33 in D. canaliculata ).

Diplommatina auriculata is very variable in shell size and rib density. Already Möllendorff (1897) described several varieties. A principal component analysis based on the five shell measurements shown in Fig. 1 as well as whorl and rib counts of 115 specimens ( Fig. 45 ) showed a broad overlap between specimens from different regions, although a vague differentiation between populations from western and eastern Java can be recognized.

There are sinistral specimens from the Botanical Garden near Cibodas ( ZMA 55000a) and Kandang Badak between Mt . Pangrango and Mt. Gede ( ZMA 68398).

Habitat. In leaf litter, on deadwood, on rocks and stones. The species can be found in shrubs, agroforest, plantations, in primary and secondary forests on calcareous and volcanic soils at 0–3000 m a.s.l.

Type material. Indonesia, Java (lectotype of D. auriculata [designated by Zilch 1953] SMF 105219; paralectotypes of D. auriculata SMF 105220/5; lectotype of D. auriculata var. abbreviata [designated by Zilch 1953] SMF 105222, paralectotypes of D. auriculata var. abbreviata SMF 105223/11; lectotype of D. auriculata var. biangulata [designated by Zilch 1953] SMF 105237, paralectotypes of D. auriculata var. biangulata SMF 105238/10). —West Java: Bogor (= Buitenzorg), 6°37'S 106°43'E (paralectotype of D. hortulana ZMH 83651/ 1).—East Java: Tengger Mountains (lectotype of D. auriculata var. gracilior [designated by Zilch 1953] SMF 105246, paralectotypes of D. auriculata var. gracilior SMF 105247/2).

Other material. Indonesia, Java (SMF 192586/4, SMF 192587/5, SMF 202838/2; RMNH 157194/1).— Banten: Lebak, Mt. Kembang, Saketeng Cave, 43 m a.s.l., 6°58'14"S 106°16'02"E (MZB 18857/6; ZMH 133044/ 1); Lebak, Mt. Kembang, 81 m a.s.l., 6°58'21"S 106°16'46"E (MZB 18856/3; ZMH 133043/4); Lebak, Cipanas, Kadir Cave, 34 m a.s.l., 6°58'41"S 106°19'18"E (MZB 18855/3; ZMH 133042/1); Lebak, Cipanas, teak plantation near Lalay Cave, 51 m a.s.l., 6°58'44"S 106°19'19"E (MZB 18861/2).—West Java: West Java (SMF 105245/1); Sukabumi, Cisolok towards the hot springs (Cipanas), 200 m a.s.l., 6°56'S 106°27'E (van Benthem Jutting 1940); Bogor, Gunung Halimun-Salak National Park, Near Pasir Ipis, 872 m a.s.l., 6°38'56"S 106°27'17"E (MZB 17978); Bogor, Gunung Halimun-Salak National Park, Pasir Ipis, Cisarua, 849 m a.s.l., 6°39'00"S 106°27'17"E (MZB 17979); Bogor, Cigudeg Village, Gudawang cave, 89 m a.s.l., 6°27'0''S 106°31'0''E (MZB 13440/1); Sukabumi, Pelabuhan Ratu, road of Cisolok-Cipanas, 6°59'S 106°33'E (SMF 105236/7); Sukabumi, Cibitung village, Cikaso waterfall 58 m a.s.l., 7°21'37"S 106°37'04"E (MZB 18859/3; ZMH 133046/4); Sukabumi, Cibitung village, Cikaso waterfall, 61 m a.s.l., 7°21'40"S 106°37'05"E (MZB 18860/9; ZMH 133047/5); Sukabumi, Cibitung village, Cikaso waterfall, 47 m a.s.l., 7°21'39"S 106°37'07"E (MZB 18862/3); Sukabumi, Cibitung village, near Cikaso waterfall, 47 m a.s.l., 7°21'40"S 106°37'09"E (MZB 18863/2); Bogor, Leuwiliang, near waterfall of Cianten (Curug Lontar), 300 m a.s.l., 6°38'13''S 106°38'18''E (SMF 202836/8; ZMA 68354/100, vouchers to van Benthem Jutting 1948); Sukabumi, Ciareuy Village, 506 m a.s.l., 7°01'32"S 106°47'59"E (MZB 18864/1); Bogor Botanical Garden 12A 12C, 6°36'S 106°48'E (ZMA 318409/2); Sukabumi, Nyalindung Village, Cipicung Cave (Buni Ayu), 755 m a.s.l, 7°02'09"S 106°54'20"E (MZB 18865/2); Sukabumi, Nyalindung Village, Bibijilan waterfall, 716 m a.s.l., 7°01'26"S 106°54'20"E (MZB 18866/2); Sukabumi, Nyalindung Village, Bibijilan waterfall, 696 m a.s.l., 7°01'27"S 106°54'21"E (MZB 18858/3; ZMH 133045/1); Sukabumi, Nyalindung Village, Buniayu, 795 m a.s.l., 7°02'11"S 106°54'25"E (MZB 18854/5; ZMH 133041/3); Sukabumi, Nyalindung Village, Buniayu, Rasamala ( Altingia ) forest, 805 m a.s.l., 7°02'10"S 106°54'34"E (MZB 18867/6); Sukabumi, Nyalindung Village, Buniayu, Rasamala ( Altingia ) forest, 777 m a.s.l., 7°02'15"S 106°54'37"E (MZB 18868/2); Sukabumi, 700 m a.s.l., 6°55'S 106°55'E (SMF 105233/21, vouchers to Paravicini 1935); Sukabumi, Siluman Cave, 789 m a.s.l., 7°01'31''S 106°55'35''E (MZB 13722/1); Sukabumi, Cibadak, Pasir Salam, 1224 m a.s.l., 7°02'S 107°02'E (BDJ); Jampang, limestone hill near Sukanegara, 1000 m a.s.l., 7°01'S 107°22'E (ZMA 68347/84, vouchers to van Benthem Jutting 1948); Bogor, Ciampea, Mt. Cibodas, 300 m a.s.l., 6°33'12''S 106°41'05''E (ZMA 68351/89, vouchers to van Benthem Jutting 1948; ZMA 317649/1); Sukabumi, Gunung Halimun-Salak National Park, Pasir Gede, 1245 m a.s.l., 6°44'17"S 106°42'53"E (MZB 17980); Bogor, Mt. Salak, near waterfall, 6°41'S 106°44'E (ZMA 68356/3); Bogor, Sukamantri, Mt. Salak, 888 m a.s.l., 6°41'S 106°45'E (BDJ/2); Bogor, Mt. Gede, 900 m a.s.l., 6°46'S 106°52'E (SMF 105240/1); Cianjur, Mt. Gede, 6°47'S 106°59'E (SMF 105232/2, 105234/2, 105244/2); Cianjur, Kandang Badak, saddle between Mt. Pangrango and Mt. Gede, 2400 m a.s.l., 6°47'S 106°58'E (ZMA 68360/98, 68398/1, vouchers to van Benthem Jutting 1948); Bogor, Mt. Pangrango, 2400 m a.s.l., 6°46'S 106°58'E (ZMA 68353/53, 68361/9, vouchers to van Benthem Jutting 1948); Bogor, Mt. Pangrango, 3000 m a.s.l., 6°46'S 106°58'E (SMF105227/11); Cianjur, Cibeureum, Mt. Gede, 1600 m a.s.l., 6°45'S 106°59'E (SMF 105235); Cianjur, Mt. Gede, near waterfalls of Cibeureum, 1700 m a.s.l., 6°45'14''S 106°59'09''E (MZB 4061/4; ZMA 33653/10, vouchers to van Benthem Jutting 1948); Cianjur, Cibeureum, 1800 m a.s.l., 6°45'14''S 106°59'09''E (MZB 4063/4); Cianjur, Mt. Gede, 2300 m a.s.l., 6°46'S 106°59'E (SMF 105226/4, 105242/5); Cianjur, Mt. Gede, Pancoran Mas near Rarahan, 1400 m a.s.l., 6°43'S 107°00'E (ZMA 68352/26, vouchers to van Benthem Jutting 1948); Puncak Pass, 1800 m a.s.l., 6°41'S 107°00'E (BDJ); Cianjur, Cibodas, Mt. Gede, 1250 m a.s.l., 6°44'34''S 107°00'39''E (RMNH 329437/2); Cianjur, Cibodas, 1400 m a.s.l., 6°45'S 107°01'E (MZB 4062/10; ZMA 68349/35, 318406/1, 318407/10, 318412/16, 318415/17); Cianjur, Cibodas, Mt. Gede, forest near Pasanggrahan, 1400 m a.s.l., 6°45'S 107°01'E (ZMA 33652/2, vouchers to van Benthem Jutting 1948); Cianjur, Cibodas, Mt. Gede, 1500 m a.s.l., 6°45'S 107°00'E (SMF 105241/10; ZMA 317645/9, 318405/2, 318416/7); Cianjur, Cibodas, Botanical Garden, 1400 m a.s.l., 6°45'S 107°00'E (ZMA 317643/2, 317644/1, 317646/7, 317648/4, 318408/4, 318410/8, 318411/1, 318413/1, 318414/7, 55000/2); West Bandung, Pangalengan, Mt. Tilu, 1900 m a.s.l., 7°06'23''S 107°23'10''E (MZB 9520/2; RMNH 44521/28; SMF 105228/2); Purwakarta, 187 m a.s.l., 6°35'S 107°25'E (RMNH 44453/6); West Bandung, Padalarang, Mt. Pawon, 700 m a.s.l., 6°49'S 107°26'E (MZB 13724/1; ZMA 68359/5, vouchers to van Benthem Jutting 1948); Bandung, Ciwidey, 1100 m a.s.l., 7°05'S 107°27'E (ZMA 68348/1, vouchers to van Benthem Jutting 1948); West Bandung, Mt. Tangkuban Prahu, 6°45'S 107°37'E (ZMA 317647/1); Garut, Sukapura, Mt. Jampang, 600 m a.s.l., 7°26'S 107°37'E (SMF 105225/12, 105239/1); Bandung, Mt. Wayang, 2000 m a.s.l., 7°13'S 107°38'E (SMF 105224/2); West Bandung, road to Lembang near Maribaya, 6°49'S 107°39'E (ZMA 68355/1); Bandung, Maribaya, 1500 m a.s.l., 6°51'S 107°42'E (ZMA 68350/2); Garut, Mt. Papandayan, 1500–1800 m a.s.l., 7°13'S 107°44'E (SMF 105243/4, 105248/1); Garut, Mt. Papandayan, forest at Tegal Primula (Tegal Panjang), 2200 m a.s.l., 7°16'S 107°44'E (ZMA 68357/2, vouchers to van Benthem Jutting 1948); Garut, Mt. Cikuray, 2500 m a.s.l., 7°19'14''S 107°51'56''E (SMF 105230/2, vouchers to Zilch 1953); Garut, Mt. Cikuray, 1900 m a.s.l., 7°18'S 107°52'E (SMF 105229/1); Garut, Mt. Cikuray, 2400 m a.s.l., 7°19'S 107°52'E (van Benthem Jutting 1948); Garut, Mt. Cikuray, 2800 m a.s.l., 7°19'S 107°52'E (SMF 105231/5, vouchers to Zilch 1953); Garut, Sukaharjo Village, Talaga Bodas, 1562 m a.s.l., 7°11'28"S 108°03'34"E (MZB 18010/5, 18012/2, 18019/1); Garut, Cinta Manik Village, Karaha Bodas, 1148 m a.s.l., 7°07'24"S 108°04'35"E (MZB 18031/19); Tasikmalaya, Linggajati Village, Mt. Galunggung, 1154 m a.s.l., 7°15'22"S 108°04'53"E (MZB 17467/1); Tasikmalaya, Karangnunggal, near Sela Dower cave, 237 m a.s.l., 7°40'00"S 108°07'00"E (MZB 18847/10; ZMH 133034/7); Tasikmalaya, Karangnunggal, surroundings of Pasir Taraje Cave, 243 m a.s.l., 7°39'58"S 108°07'01"E (MZB 18848/5; ZMH 133035/3); Tasikmalaya, Karangnunggal, surroundings of Liang cai, 237 m a.s.l., 7°40'01"S 108°07'03"E (MZB 18846/43; ZMH 133033/10); Tasikmalaya, Karangnunggal, mouth of Bojong Cave, 240 m a.s.l., 7°40'12"S 108°07'12"E (MZB 18845/8; ZMH 133032/4); Tasikmalaya, Setiawaras Village, near Sambawa Cave (Ranggawulung Cave), 361 m a.s.l., 7°35'02"S 108°10'57"E (MZB 17774/7); Ciamis, Sukamaju Village, Mt. Sawal Wildlife Reserve, 1017 m a.s.l., 7°12'S 108°15'E (MZB 16405/1, 16453/1); Majalengka, Argamukti, Mt. Ciremai National Park, 6°54'S 108°22'E (MZB 13258/69, vouchers to Heryanto 2008, 2009 [as D. auriculata ]; MZB 17975/2, vouchers to Heryanto 2008, 2009 [as D. cyclostoma ]); Kuningan, Kiara Bolong, Mt. Ciremai, 2320 m a.s.l., 6°55'S 108°24'E (BDJ/3); Pangandaran, 28 m a.s.l., 7°42'23''S 108°39'22''E (MZB 10033/2); Pangandaran, 7°41'S 108°40'E (JV 4106/8); Pangandaran, Rengganis Cave, 41 m a.s.l., 7°43'S 108°40'E (BDJ/1)—Central Java: Cilacap,west part of Nusa Kambangan Island, Kali Jati, 1 m a.s.l., 7°43'S 108°48'E (BDJ/2); Cilacap, Nusa Kambangan, Ratu Batu cave, 39 m a.s.l., 7°44'S 108°54'E (MZB 9481/1; RMNH 44452/22, vouchers to Vermeulen & Whitten 1998); Tegal, Kedawung Village, Jurangmangu, Mt. Slamet, 1350 m a.s.l., 7°12'S 109°12'E (BDJ/7); Purwokerto, Kali Pagu Village, Mt. Slamet, 638 m a.s.l., 7°19'S 109°12'E (MZB 15396/4, 15405/3, 15411/4, 15415/22, 15419/15, 15423/27, 15428/16, 15434/7, 15464/4, 15492/1, 15499/4, 15529/3, vouchers to Heryanto 2012); Mt. Slamet, Kali Mertelu 107 m a.s.l. (MZB 15387/42, vouchers to Heryanto 2012); Kebumen, Karang Bolong, Petruk Cave, 78 m a.s.l., 7°42'17"S 109°23'59"E (MZB 18849/5; ZMH 133036/3); Kebumen, Karang Bolong, Petruk Cave, 103 m a.s.l., 7°42'18"S 109°24'02"E (MZB 18869/2); Kebumen, Karang Bolong, wall and surroundings of Petruk Cave, 128 m a.s.l., 7°42'20"S 109°24'02"E (MZB 18851/13; ZMH 133038/4); Kebumen, Karang Bolong, Petruk Cave, 138 m a.s.l., 7°42'18"S 109°24'05"E (MZB 18850/7; ZMH 133037/4); Kebumen, Karang Bolong, inside Petruk Cave, (MZB 18852/3; ZMH 133039/1); Kebumen, Karang Bolong, Adiraja Hill, 170 m a.s.l., 7°42'50"S 109°24'05"E (MZB 18870/4); Kebumen, Karang Bolong, Adiraja Hill, 172 m a.s.l., 7°42'47"S 109°24'05"E (MZB 18853/3; ZMH 133040/1); Kebumen, west of Karang Bolong Mts, Petruk Cave, 352 m a.s.l., 7°43'S 109°26'E (RMNH 44454/7); Purworejo, Donorejo village, near Seplawan Cave, 700 m a.s.l., 7°45'52''S 110°5'46''E (MZB 17754/4).—Yogyakarta: Kulonprogo, Girimulyo, Sibolong Hill, 721 m a.s.l., 7°45'0''S 110°7'0"E (MZB 17757/3); Kulonprogo, Sabrang Kidul Village, 730 m a.s.l., 7°44'38''S 110°7'55''E (MZB 17668/ 2); Kulonprogo, Sibolong Village, 716 m a.s.l., 7°44'44''S 110°8'2''E (MZB 17704/4).—East Java: Cemorosewu near Sarangan, 1875 m a.s.l., 7°39'52''S 111°11'29''E ( Rensch 1934); Magetan, Kali Jumok, Mt. Lawu, 1787 m a.s.l., 7°39'S 111°13'E (ZMB 76098/3; Rensch 1934); Magetan, Sarangan Village, 1300 m a.s.l., 7°40'28''S 111°13'19''E (MZB 14170/2); Malang, Sempu Island, Baru-baru Hill, 29 m a.s.l., 8°26'52"S 112°41'17"E (MZB 18871/2); Malang Sempu Island, Baru-baru crossroad, 38 m a.s.l., 8°26'53"S 112°41'24"E (MZB 18843/3; ZMH 133031/1); Malang, Sempu Island, Wall of Kelabang Cave, 44 m a.s.l., 8°26'58"S 112°41'28"E (MZB 18844/5); Malang, Sempu Island, Jembatan Anakan Hill, 46 m a.s.l., 8°27'05"S 112°41'28"E (MZB 18872/1); Pasuruan, Nongkojajar, Tengger Mts., 1250 m a.s.l., 7°54'S 112°50'E (van Benthem Jutting 1948); Tengger Mts. (SMF 105247/2); Probolinggo, Taman Hidup, 1900 m a.s.l., 7°59'S 113°32'E (RMNH 44457/7); Surabaya, 7°16'S 112°43'E (SMF 105221/5, vouchers to Zilch 1953); Probolinggo, Bromo, near Mt. Argopuro, 1200 m a.s.l., 7°56'S 112°60'E (RMNH 44456/2); Situbondo, Kendeng III, Mt. Ijen, 1400 m a.s.l., 7°59'S 114°13'E (ZMA 68358/1, vouchers to van Benthem Jutting 1948); Banyuwangi, Alas Purwo National Park, near Sadengan, 29 m a.s.l., 8°39'04"S 114°22'20"E (MZB 18873/3); Banyuwangi, Alas Purwo National Park, on the way to Padepokan Cave, 93 m a.s.l., 8°40'03"S 114°23'07"E (MZB 18875/1); Banyuwangi, Alas Purwo National Park, on the way to Gua Mayangkoro, 120 m a.s.l., 8°40'12"S 114°23'09"E (MZB 18874/1).

Distribution ( Fig. 17 ). Diplommatina auriculata is known from Java and Bali ( Vermeulen & Whitten 1998).

D D3W H da ha W D/H D/da H/ha da/ha rpw rbw

Diplommatina planicollis , n=20

min 2.2 0.85 4.5 1.4 1.0 7 0.47 1.44 2.82 0.91 56 38 max 2.6 5 1.0 5 5.1 1.7 1.7 5 7.7 5 0.5 6 1.6 7 4.7 5 1.1 1 8 4 5 5 mean 2.5 0.9 4.8 1.6 1.5 7.2 0.51 1.58 3.19 1.02 69.9 48.1 SD 0.1 0.1 0.2 0.4 0.2 0.3 0.02 0.06 0.39 0.06 23.9 25.2 Diplommatina sulcicollis , n=20

min 1.5 1.05 2.5 0.95 0.95 5 0.55 1.30 2.39 0.95 88 51 max 1.6 5 1.4 2.8 1.1 5 1.1 5 5.2 5 0.6 0 1.6 0 2.8 0 1.1 5 1 1 3 7 5 mean 1.6 1.2 2.7 1.1 1.0 5.0 0.57 1.44 2.63 1.04 98.3 62.6 SD 0.0 0.1 0.1 0.1 0.1 0.1 0.02 0.09 0.12 0.06 24.7 6.2 Diplommatina halimunensis , n=3

min 1.4 5 0.8 2.4 5 0.8 0.7 5.5 0.5 4 1.2 4 2.4 3 1.1 9 1 8 1 8 max 1.5 5 0.9 2.8 1.2 5 1.0 5 5.7 5 0.6 1 1.8 1 3.6 3 1.2 0 2 3 2 4 mean 1.5 0.9 2.6 1.1 0.9 5.6 0.58 1.43 2.95 1.19 21 21.7 SD 0.1 0.1 0.2 0.2 0.2 0.1 0.04 0.33 0.61 0.01 2.7 3.2 D D3W H da ha W D/H D/da H/ha da/ha rpw rbw

Diplommatina heryantoi , n=20

min 1.25 1.05 2.1 0.8 0.75 5 0.53 1.48 2.63 1.00 50 35 max 1.4 1.2 2.4 0.9 0.9 6 0.62 1.67 3.07 1.17 76 54 mean 1.3 1.1 2.3 0.8 0.8 5.7 0.58 1.57 2.87 1.06 58.0 42.5 SD 0.1 0.0 0.1 0.0 0.0 0.2 0.02 0.05 0.12 0.05 6.5 4.9 Diplommatina diplostoma , n=25

min 1.6 1.2 2.6 1.0 0.8 5.25 0.58 1.46 2.40 1.00 24 22 max 2.6 1.6 4.0 1.6 1.6 6.25 0.70 1.83 3.44 1.31 87 53 mean 1.8 1.4 2.9 1.1 1.0 5.7 0.63 1.62 2.89 1.12 69.0 43.8 SD 0.3 0.1 0.4 0.2 0.2 0.2 0.02 0.08 0.26 0.08 17.4 7.6 Diplommatina perpusilla , n=40

min 0.85 0.55 1.6 0.6 0.55 5 0.48 1.27 2.43 1.00 36 21 max 1.1 0.85 2.05 0.8 0.7 6 0.66 1.62 3.36 1.36 61 34 mean 1.0 0.7 1.7 0.7 0.6 5.6 0.57 1.40 2.79 1.13 45.3 27.4 SD 0.1 0.1 0.1 0.0 0.0 0.3 0.03 0.08 0.19 0.08 9.6 8.0 Diplommatina ristiae , n=11

min 0.95 0.65 1.65 0.6 0.6 5.25 0.53 1.40 2.71 1.00 22 12 max 1.25 0.75 2.0 0.75 0.7 6 0.68 1.67 3.08 1.17 31 16 mean 1.1 0.7 1.8 0.7 0.6 5.9 0.58 1.52 2.90 1.10 25.8 14.0 SD 0.1 0.0 0.1 0.1 0.0 0.3 0.04 0.08 0.10 0.06 4.1 2.8 Diplommatina calcarata , n=26

min 1.3 0.65 2.1 0.8 0.8 5.5 0.57 1.53 2.35 0.91 15 13 max 1.7 0.9 2.65 1.1 0.95 6.75 0.74 1.78 2.94 1.17 28 25 mean 1.5 0.8 2.3 0.9 0.9 6.0 0.63 1.64 2.66 1.03 20.1 18.8 SD 0.1 0.1 0.2 0.1 0.0 0.4 0.04 0.07 0.17 0.06 4.4 3.4 Diplommatina tetragonostoma , n=6

min 1.4 0.7 2.5 0.85 0.85 6.5 0.52 1.40 2.61 1.00 31 31 max 1.65 0.85 2.9 1.1 1.1 6.75 0.60 1.76 3.15 1.17 39 43 mean 1.5 0.8 2.7 1.0 0.9 6.6 0.57 1.56 2.91 1.07 33.8 36.2 SD 0.1 0.1 0.1 0.1 0.1 0.1 0.03 0.13 0.20 0.07 3.0 5.0 Diplommatina baliana , n=70

min 1.05 0.65 2.2 0.75 0.7 5.75 0.39 1.17 2.67 0.88 36 20 max 1.5 0.9 2.85 0.95 0.95 6.75 0.56 1.77 3.53 1.20 60 37 mean 1.3 0.8 2.6 0.9 0.8 6.1 0.50 1.44 3.06 1.06 48.8 27.4 SD 0.1 0.1 0.2 0.2 0.1 0.3 0.03 0.10 0.20 0.06 5.1 3.4 Diplommatina nevilli , n=28

min 1.05 0.55 2.25 0.70 0.5 6.25 0.41 1.30 2.94 0.94 23 17 max 1.5 0.8 2.9 1.0 0.85 7.5 0.52 1.60 4.60 1.50 32 25 mean 1.1 0.6 2.5 0.8 0.7 6.7 0.45 1.42 3.47 1.11 26.9 20.8 SD 0.1 0.1 0.1 0.1 0.1 0.2 0.02 0.07 0.26 0.08 2.2 1.5 D D3W H da ha W D/H D/da H/ha da/ha rpw rbw

Diplommatina canaliculata , n=20

min 1.2 0.55 3.05 0.9 0.9 7 0.40 1.16 3.00 0.97 24 16 max 1.7 0.75 3.8 1.3 1.15 7.75 0.49 1.45 3.58 1.25 33 26 mean 1.6 0.6 3.3 1.2 1.0 7.5 0.47 1.34 3.23 1.13 26.6 22.0 SD 0.1 0.0 0.2 0.1 0.1 0.2 0.02 0.07 0.15 0.07 2.3 1.9 Diplommatina auriculata , n=115

min 1.45 0.6 2.9 1.0 0.85 6.5 0.45 1.24 2.61 0.88 28 25 max 2.2 1.05 4.45 1.65 1.45 8 0.58 1.71 3.82 1.36 92 77 mean 1.7 0.7 3.4 1.2 1.1 7.2 0.51 1.46 3.17 1.10 52.6 39.8 SD 0.2 0.1 0.4 0.1 0.1 0.4 0.03 0.08 0.24 0.09 11.2 8.8

Figures

FIGURE 45. Principal component analySiS baSed on fiVe meaSurementS aS well aS whorl and rib countS of 115 SpecimenS of Diplommatina auriculata Möllendorff, 1897.

FIGURES 1 – 3. Shell terminology. 1. MeaSurementS. AbbreViationS: D, Shell diameter; D 3 W, diameter of the third whorl; da, diameter of the aperture; H, Shell height; ha, height of the aperture. 2. Terminology of the inner lamellae and foldS in the laSt whorl. AbbreViationS: cl, columellar lamella, pl, parietal lamella; rpf, radial palatal fold; Spf, Spiral palatal fold. 3. PoSitional termS. Shell Shown from the baSe. AbbreViationS: d, dorSally; dll, dorSolaterally left; dlr, dorSolaterally right; ll, laterally left; lr, laterally right; V, Ventrally; Vll, Ventrolaterally left; Vlr, Ventrolaterally right.

FIGURES 4 – 8. Opercula of Diplommatinidae. 4. Diplommatina javana Möllendorff, 1897, outer and inner Surface of opercula, WeSt JaVa: Majalengka, Argamukti, Mt. Ciremai National Park (MZB 13261). 5. Diplommatina auriculata Möllendorff, 1897, inner Surface of operculum, WeSt JaVa: Majalengka, Argamukti, Mt. Ciremai National Park (MZB 13258). 6. Palaina gedeana Möllendorff, 1897, outer and inner Surface of opercula, WeSt JaVa: Majalengka, Argamukti, Mt. Ciremai National Park (MZB 13244). 7. Palaina nubigena Möllendorff, 1897, outer Surface of operculum, WeSt JaVa: Cianjur, Kandang Badak (ZMA 68468). 8. Arinia yanseni n. sp., inner Surface of operculum, EaSt JaVa: Rembang, Sumber PacuSen (MZB 18882). Scale bar = 0.07 mm for FigS 3 – 6, 0.1 mm for Fig. 7.

FIGURES 15 – 19. DiStribution of Diplommatinidae in JaVa. 15. Diplommatina baliana Fulton, 1899. 16. Diplommatina diplostoma RenSch, 1931, Diplommatina nevilli (CroSSe, 1879) and Diplommatina canaliculata Möllendorff, 1887. 17. Diplommatina auriculata Möllendorff, 1897. 18. Palaina gedeana Möllendorff, 1897, Palaina nubigena Möllendorff, 1897 and Arinia yanseni n. sp. 19. Opisthostoma javanicum Van Benthem Jutting, 1932, Opisthostoma uranoscopium Van Benthem Jutting, 1932 and Opisthostoma sp.

FIGURES 41 – 42. Diplommatina auriculata Möllendorff, 1897 from JaVa. 41. JaVa (lectotype of D. auriculata SMF 105219). 42. JaVa (lectotype of D. auriculata Var. abbreviata SMF 105222). Scale bar = 1.0 mm.

FIGURES 43 – 44. Diplommatina auriculata Möllendorff, 1897 from JaVa. 43. JaVa (lectotype of D. auriculata Var. biangulata SMF 105237). 44. EaSt JaVa: Tengger MountainS (lectotype of D. auriculata Var. gracilior SMF 105246). Scale bar = 1.0 mm.