Diplommatina perpusilla Möllendorff, 1897

Hausdorf, Bernhard, 2017, Revision of the Diplommatinidae (Gastropoda: Cyclophoroidea) from Java, Zootaxa 4312 (2), pp. 201-245 : 216-218

Diplommatina perpusilla Möllendorff, 1897

( Figs 12 , 29 ; Table 1)

Diplommatina (Sinica) perpusilla Möllendorff, 1897: 96 (JaVa) . Kobelt 1902: 470. Zilch 1953: 35, pl. 11 fig. 169.

Diplommatina perpusilla — Van Benthem Jutting 1948: 591, fig. 48. Heryanto 2009: 332 (partly?). Heryanto 2011: 54, fig.

Diplommatina cyclostoma —Heryanto 2008: 362 (partly, not Möllendorff 1897). Heryanto 2009: 332 (partly, not Möllendorff 1897).

Diplommatina duplicilabra —Heryanto 2008: 362 (partly, not Van Benthem Jutting 1948). Heryanto 2009: 332 (partly, not Van Benthem Jutting 1948).

Palaina gedeana —Heryanto 2008: 362 (partly, non Möllendorff, 1897). Heryanto 2009: 332 (partly, non Möllendorff, 1897).

Diagnosis. Diplommatina perpusilla is characterized by its minute-sized, distinctly ribbed shell, the ventrolaterally right or ventral constriction of the last whorl and the ventrally or ventrolaterally left position of the spiral palatal fold.

Shell ( Fig. 29 ; Table 1). Dextral; spindle-shaped with conical early whorls and a narrowed last whorl; with 5– 6 whorls separated by a deep suture; protoconch smooth; teleoconch with slightly oblique ribs (36–61 on the penultimate whorl; fewer on the last whorl); with fine spiral striae between the ribs; whitish or corneous; subtranslucent; last whorl rounded, with an indistinct ventrolaterally right or ventral constriction; aperture almost circular, slightly angular at the transition from columellar to basal side; upper insertion of the peristome ascending towards aperture; insertions of the peristome connected by an indistinct callus; peristome expanded, thickened, doubled; aperture with a prominent columellar lamella; with a parietal lamella ventrolaterally left inside the last whorl; with a short or long spiral palatal fold in the middle of the whorl ventrolaterally left or ventrally inside the last whorl, and with a radial palatal fold ventrolaterally right or ventrally close to the suture; umbilicus closed; operculum thin, corneous, circular, paucispiral. Shell height 1.6–2.05 mm, shell diameter 0.85–1.1 mm.

Remarks. Diplommatina perpusilla differs from the probably most closely related D. ristiae n. sp. in the distinct ribbing, the ventrolaterally right or ventral constriction of the last whorl and the ventral or ventrolateral left position of the spiral palatal fold.

Diplommatina perpusilla is variable with regard to the palatal folds. Usually there is a long spiral palatal fold ventrally inside the last whorl and the constriction of the last whorl and the radial palatal fold are ventrolaterally right. In some specimens or populations (e.g. from Cibodas, Cibeureum, Talaga Bodas, Karaha Bodas) the armature is shifted towards the aperture. The constriction of the last whorl and the radial palatal fold are ventral and the spiral palatal fold is short and situated ventrolaterally left. Moreover, the peristome is often less distinctly expanded in such specimens. There are also intermediate specimens and specimens with rudimentary or even missing palatal folds (e.g. from Mt. Sawal Wildlife Reserve).

Habitat. In leaf litter and on deadwood in agroforest, plantation and secondary forest on calcareous and volcanic soils at 43–2800 m a.s.l.

Type material. Indonesia, Java (lectotype of D. perpusilla [designated by Zilch 1953] SMF 105210; paralectotypes of D. perpusilla SMF 105211/4).

Other material. Indonesia, Java, Banten: Lebak, Mt. Kembang, Saketeng Cave, 43 m a.s.l., 6°58'14"S 106°16'02"E (MZB 18840/1).—West Java: Sukabumi, Ciareuy, Jampang Tengah, 573 m a.s.l., 7°01'49"S 106°48'26"E (MZB 18841/1); Sukabumi, Nyalindung, Bibijilan waterfall, 716 m a.s.l., 7°01'26"S 106°54'20"E (MZB 18842/1); Cianjur, Mt. Gede, 1800 m a.s.l., 6°45'S 106°59'E (SMF 105213/3); Cianjur, Mt. Gede, 2000 m a.s.l., 6°46'S 106°59'E (ZMA 33698/3, vouchers to van Benthem Jutting 1948); Cianjur, along the road between Cibeureum and Kandang Badak, 2000 m a.s.l., 6°46'S 106°59'E (ZMA 68396/2, vouchers to van Benthem Jutting 1948); Cianjur, Cibodas, Kandang Badak, 2000 m a.s.l., 6°46'S 106°59'E (MZB 4064/2); Cianjur, Mt. Gedeh, 2000 m a.s.l., 6°46'S 106°59'E (MZB 4067/1); Cianjur, Cibodas, Botanical Garden, 1400 m a.s.l., 6°45'S 107°00'E (ZMA 317656/1); Cianjur, Cibodas, along path behind Pesanggrahan, 1500 m a.s.l., 6°45'S 107°00'E (RMNH 329448/1; ZMA 317655/4, 318420/2); Cianjur, Cibodas, 1400 m a.s.l., 6°45'S 107°01'E (MZB 4065/2); Cianjur, Cibeureum, 1400 m a.s.l., 6°45'S 107°01'E (MZB 4066/1); West Bandung, Mt. Tilu, 1900 m a.s.l., 7°06'S 107°23'E (SMF 105250/2, vouchers to Zilch 1953); Garut, Mt. Cikuray, 2800 m a.s.l., 7°19'S 107°52'E (SMF 105212/1, voucher to Zilch 1953); Garut, Sukaharjo Village, Talaga Bodas, 1562 m a.s.l., 7°11'28"S 108°03'34"E (MZB 18013/5); Garut, Cinta Manik Village, Karaha Bodas, 1148 m a.s.l., 7°07'24"S 108°04'35"E (MZB 18032/2, 18033/1); Ciamis, Sukamaju Village, Mt. Sawal Wildlife Reserve at the riverbanks, Citanduy Watershed, 877 m a.s.l., 7°13'S 108°14'E (MZB 16419/1, 16420/1, 16425/8, 16445/5); Ciamis, Sukamaju Village, Mt. Sawal Wildlife Reserve, Cihaurbeuti District, 1008 m a.s.l., 7°12'51"S 108°15'32"E (MZB 16411/3); Ciamis, Sukamaju Village, Mt. Sawal Wildlife Reserve, 1050 m a.s.l., 7°12'S 108°15'E (MZB 16406/6, 16456/3, 16483/1); Citanduy Watershed, Mt. Sawal Wildlife Reserve, Kampung Cikujang Girang, Sukamaju Village, Cihaurbeuti District, Ciamis, 917 m a.s.l., 7°13'16"S 108°15'20"E (MZB 16470/1); Ciamis, Sukamaju Village, Mt. Sawal Wildlife Reserve, 1017 m a.s.l., 7°12'28"S 108°15'21"E (MZB 16451/2); Ciamis, Sukamaju Village, Mt. Sawal Wildlife Reserve, 1092 m a.s.l., 7°12'20"S 108°15'22"E (MZB 16516/10); Ciamis, Sukamaju Village, Mt. Sawal Wildlife Reserve, 1138 m a.s.l., 7°12'15"S 108°15'22"E (MZB 16455/1); Ciamis, Sukamaju Village, Mt. Sawal Wildlife Reserve, 913 m a.s.l., 7°13'19"S 108°15'24"E (MZB 16472/3); Ciamis, Sukamaju Village, Mt. Sawal Wildlife Reserve, 910 m a.s.l., 7°13'21"S 108°15'25"E (MZB 16480/5); Ciamis, Sukamaju Village, Mt. Sawal Wildlife Reserve, 963 m a.s.l., 7°13'12"S 108°15'26"E (MZB 16435/1); Ciamis, Sukamaju Village, Mt. Sawal Wildlife Reserve, 909 m a.s.l., 7°13'11"S 108°15'32"E (MZB 16434/5); Ciamis, Sukamaju Village, Mt. Sawal Wildlife Reserve, 928 m a.s.l., 7°13'00"S 108°15'33"E (MZB 16513/1); Ciamis, Sukamaju Village, Mt. Sawal Wildlife Reserve, 1075 m a.s.l., 7°12'37"S 108°15'36"E (MZB 16499/3); Ciamis, Sukamaju Village, Mt. Sawal Wildlife Reserve, 1031 m a.s.l., 7°13'41"S 108°15'36"E (MZB 16505/4); Ciamis, Sukamaju Village, Mt. Sawal Wildlife Reserve, 1062 m a.s.l., 7°12'31"S 108°15'38"E (MZB 16504/5); Ciamis, Sukamaju Village, Mt. Sawal Wildlife Reserve, 1004 m a.s.l., 7°12'49"S 108°15'38"E (MZB 16498/3); Majalengka, Argamukti, Mt. Ciremai National Park, 6°54'S 108°22'E (MZB 13262/8, vouchers to Heryanto 2008, 2009 [as D. duplicilabra ]; MZB 13263/21, vouchers to Heryanto 2008, 2009 [as D. cyclostoma ]); Majalengka, Argamukti, Mt. Ciremai National Park, 1735 m a.s.l., 6°54'S 108°22'E (MZB 17972/9, vouchers to Heryanto 2008 [as P. gedeana ]); Kuningan, Palutungan, 6°55'S 108°24'E (MZB 18901/2, 18906/3, vouchers to Heryanto 2009); Kuningan, Palutungan, 1835 m a.s.l., 06°55'10"S 108°23'00"E (MZB 18903/2); Kuningan, Palutungan, 1875 m a.s.l., 06°55'20"S 108°24'40"E (MZB 14841/2, 18902/1, vouchers to Heryanto 2009); Kuningan, Palutungan, 1935 m a.s.l., 06°55'15"S 108°24'40"E (MZB 14845/1, voucher to Heryanto 2009); Mt. Ciremai, 1500 m a.s.l., 6°53'S 108°26'E (van Benthem Jutting 1948).

Distribution ( Fig. 12 ). Diplommatina perpusilla is endemic to Banten and West Java.

Figures

FIGURES 10 – 14. DiStribution of Diplommatinidae in JaVa. 10. Diplommatina planicollis Möllendorff, 1897, Diplommatina heteroglypha Van Benthem Jutting, 1948 and Diplommatina javana Möllendorff, 1897. 11. Diplommatina halimunensis n. sp., Diplommatina kakenca n. sp. and Diplommatina heryantoi n. sp. 12. Diplommatina perpusilla Möllendorff, 1897 and Diplommatina ristiae n. sp. 13. Diplommatina sulcicollis Möllendorff, 1897 and Diplommatina cyclostoma Möllendorff, 1897. 14. Diplommatina calcarata Möllendorff, 1897 and Diplommatina tetragonostoma Möllendorff, 1897.

FIGURES 29 – 32. Diplommatina Spp. from JaVa. 29. Diplommatina perpusilla Möllendorff, 1897, JaVa (lectotype SMF 105210). 30. Diplommatina ristiae n. sp., Central JaVa: Donorejo Village, near Seplawan CaVe, KaligeSing, Purworejo (holotype MZB 18886). 31. Diplommatina calcarata Möllendorff, 1897, JaVa (lectotype SMF 105207). 32. Diplommatina tetragonostoma Möllendorff, 1897, WeSt JaVa: Bandung, Mt. Wayang (holotype SMF 105199). Scale bar = 1.0 mm.