Palaina nubigena Möllendorff, 1897

Hausdorf, Bernhard, 2017, Revision of the Diplommatinidae (Gastropoda: Cyclophoroidea) from Java, Zootaxa 4312 (2), pp. 201-245 : 237-238

Palaina nubigena Möllendorff, 1897

( Figs 7 , 18 , 47 ; Table 2)

Palaina nubigena Möllendorff, 1897: 94 (JaVa; more exactly: Cikuray, See Zilch 1953). Van Benthem Jutting 1948: 586, fig. 42. Heryanto 2008: 362 (partly). Heryanto 2009: 332 (partly).

Palaina (Palaina) nubigena — Kobelt 1902: 402. Zilch 1953: 12, pl. 5, fig. 62.

Diagnosis. Palaina nubigena is characterized by an ovoid, closely ribbed, sinistral shell and an operculum with an indistinct spiral ridge on the outer surface.

Shell ( Figs 7 , 47 ; Table 2). Sinistral; ovoid with conical early whorls; with 5–5.5 whorls separated by a deep suture; protoconch smooth; teleoconch with oblique ribs (80–104 on the penultimate whorl; fewer on the last whorl); with indistinct spiral striae between the ribs; whitish; subtranslucent; last whorl rounded, without distinct constriction or protrusion; aperture almost circular; upper insertion of the peristome slightly ascending towards aperture; insertions of the peristome connected by a distinct callus; peristome hardly expanded, slightly thickened, doubled at the columellar margin; without lamellae or folds; rimate or umbilicus closed; operculum thin, corneous, circular, multispiral, with an indistinct spiral ridge on the outer surface. Shell height 2.55–3.0 mm, shell diameter 1.5–2.0 mm.

Habitat. In leaf litter in forests on volcanic soils at 2300–2800 m a.s.l.

Type material. Indonesia, Java, West Java: Cikuray (= Tjikorai), 7°19'S 107°52'E (lectotype of P. nubigena [designated by Zilch 1953] SMF 105095; paralectotypes of P. nubigena SMF 105096/10).

Other material. Indonesia, Java (SMF 192462/4).—West Java: Cianjur, Mt. Pangrango, 2400 m a.s.l., 6°46'S 106°58'E (ZMA 68465/14, 68466/1, voucher to van Benthem Jutting 1948); Cianjur, Kandang Badak, saddle between Mt.Gede and Mt.Pangrango, 2400 m a.s.l., 6°47'S 106°58'E (ZMA 68468/66, voucher to van Benthem Jutting 1948); Cianjur, Mt. Gede (SMF 105099/2); Cianjur, Mt. Gede, 2300–2500 m a.s.l., 6°46'S 106°59'E (SMF 105098/8, voucher to Zilch 1953); Cianjur, Mt. Gede, 2800 m a.s.l., 6°48'S 106°59'E (van Benthem Jutting 1948); Garut, top of Mt. Cikuray (= Tjikorai), 2600 m a.s.l., 7°19'S 107°52'E (SMF 105097/4, vouchers to Zilch 1953).

Distribution ( Fig. 18 ). Palaina nubigena is endemic to West Java, where it has been found so far in the Mt. Gede - Mt. Pangrango range and on Mt. Cikuray.

Figures

FIGURES 4 – 8. Opercula of Diplommatinidae. 4. Diplommatina javana Möllendorff, 1897, outer and inner Surface of opercula, WeSt JaVa: Majalengka, Argamukti, Mt. Ciremai National Park (MZB 13261). 5. Diplommatina auriculata Möllendorff, 1897, inner Surface of operculum, WeSt JaVa: Majalengka, Argamukti, Mt. Ciremai National Park (MZB 13258). 6. Palaina gedeana Möllendorff, 1897, outer and inner Surface of opercula, WeSt JaVa: Majalengka, Argamukti, Mt. Ciremai National Park (MZB 13244). 7. Palaina nubigena Möllendorff, 1897, outer Surface of operculum, WeSt JaVa: Cianjur, Kandang Badak (ZMA 68468). 8. Arinia yanseni n. sp., inner Surface of operculum, EaSt JaVa: Rembang, Sumber PacuSen (MZB 18882). Scale bar = 0.07 mm for FigS 3 – 6, 0.1 mm for Fig. 7.

FIGURES 15 – 19. DiStribution of Diplommatinidae in JaVa. 15. Diplommatina baliana Fulton, 1899. 16. Diplommatina diplostoma RenSch, 1931, Diplommatina nevilli (CroSSe, 1879) and Diplommatina canaliculata Möllendorff, 1887. 17. Diplommatina auriculata Möllendorff, 1897. 18. Palaina gedeana Möllendorff, 1897, Palaina nubigena Möllendorff, 1897 and Arinia yanseni n. sp. 19. Opisthostoma javanicum Van Benthem Jutting, 1932, Opisthostoma uranoscopium Van Benthem Jutting, 1932 and Opisthostoma sp.

FIGURES 46 – 48. Diplommatinidae from JaVa. 46. Palaina gedeana Möllendorff, 1897, JaVa (lectotype SMF 105100). 47. Palaina nubigena Möllendorff, 1897, WeSt JaVa: Cikuray (lectotype SMF 105095). 48. Arinia yanseni n. sp., EaSt JaVa: Rembang, Sumber PacuSen (holotype MZB 18893). Scale bar = 1.0 mm.