Palaina gedeana Möllendorff, 1897

Hausdorf, Bernhard, 2017, Revision of the Diplommatinidae (Gastropoda: Cyclophoroidea) from Java, Zootaxa 4312 (2), pp. 201-245 : 236

Palaina gedeana Möllendorff, 1897

( Figs 6 , 18 , 46 ; Table 2)

Palaina gedeana Möllendorff, 1897: 94 (JaVa; more exactly: Mt. Gede, See Kobelt 1902). Van Benthem Jutting 1948: 585, fig. 40–41. Vermeulen & Whitten 1998: 51, 139, fig. 32. Heryanto 2008: 362 (partly). Heryanto 2009: 332 (partly). Heryanto 2011: 56, fig.

Palaina (Palaina) gedeana — Kobelt 1902: 399. Zilch 1953: 10, pl. 5, fig. 61.

Diagnosis. Palaina gedeana is characterized by a cylindrical, closely ribbed, dextral shell with an operculum with a high spiral ridge on the outer surface.

Shell ( Fig. 6 , 46 ; Table 2). Dextral; cylindrical with conical early whorls; with 5–5.5 whorls separated by a deep suture; protoconch smooth; teleoconch with oblique ribs (58–79 on penultimate whorl; fewer on the last whorl); with or without distinct spiral striae between the ribs; whitish; subtranslucent; last whorl rounded, with an indistinct ventral constriction, with a ventrolateral left protrusion; aperture almost circular; upper insertion of the peristome slightly ascending towards aperture; insertions of the peristome connected by a distinct callus; peristome hardly expanded, slightly thickened, doubled or not doubled; without lamellae or folds; rimate; operculum thin, corneous, circular, multispiral, with a spiral ridge on the outer surface and an arcuate ridge on the inner surface. Shell height 2.1–2.9 mm, shell diameter 1.15–1.55 mm.

Remarks. The protrusion of the last whorl is more distinct in specimens from East Java.

Habitat. In leaf litter in primary and secondary forests on volcanic soils at 1400–3000 m a.s.l.

Type material. Indonesia, Java (lectotype of P. gedeana [designated by Zilch 1953] SMF 105100; paralectotypes of P. gedeana (SMF 105101/12, SMF 105103/8, SMF 105104/6, SMF 105106/12).

Other material. Indonesia, Java, West Java: Cianjur, Mt. Pangrango, 2400 m a.s.l., 6°46'S 106°58'E (ZMA 33803/100, 68453/44); Cianjur, Mt. Pangrango, 2900 m a.s.l., 6°46'S 106°58'E (ZMA 68459/7); Cianjur, Kandang Badak, saddle between Mt.Gede and Mt.Pangrango, 2400 m a.s.l., 6°47'S 106°58'E (SMF 201234 /5; ZMA 68455/ 100, vouchers to van Benthem Jutting 1948); Mt. Gede (SMF 192461/4); Cianjur, Mt. Gede, 2500 m a.s.l., 6°47''S 106°58'E (SMF 105105/6, vouchers to Zilch 1953); Cianjur, Mt. Gede, 2900 m a.s.l., 6°47'S 106°59'E (van Benthem Jutting 1948); Cianjur, Mt. Gede, 1800 m a.s.l., 6°45'S 106°59'E (SMF 105102/7, vouchers to Zilch 1953); Cianjur, Cibodas, 1400 m a.s.l., 6°45'S 107°01'E (MZB 4068/2); Garut, Cikuray (SMF 105215a/15); Majalengka, Argamukti, Mt. Ciremai National Park, 1735 m a.s.l., 6°54'S 108°22'E (MZB 13244/16, vouchers to Heryanto 2008).—East Java: Probolinggo, Taman Hidup, 1900 m a.s.l., 7°59'S 113°32'E (RMNH 44741/48); Probolinggo, Taman Hidup, 2100 m a.s.l., 7°59'S 113°32'E (RMNH 44742/2), Bondowoso, Mt. Ijen, Merapi Ungup-ungup, 1850 m a.s.l., 8°05'S 114°14'E (ZMA 68452/8, ZMA 68458/12, vouchers to van Benthem Jutting 1948); Bondowoso, game reserve Ijen Crater-Merapi, 1600–1700 m a.s.l., 8°05'S 114°14'E (ZMA 68454/2, 68456/ 5, 68457/24, vouchers to van Benthem Jutting 1948).

Distribution ( Fig. 18 ). Palaina gedeana is known from Java and Bali ( Vermeulen & Whitten 1998).

Figures

FIGURES 4 – 8. Opercula of Diplommatinidae. 4. Diplommatina javana Möllendorff, 1897, outer and inner Surface of opercula, WeSt JaVa: Majalengka, Argamukti, Mt. Ciremai National Park (MZB 13261). 5. Diplommatina auriculata Möllendorff, 1897, inner Surface of operculum, WeSt JaVa: Majalengka, Argamukti, Mt. Ciremai National Park (MZB 13258). 6. Palaina gedeana Möllendorff, 1897, outer and inner Surface of opercula, WeSt JaVa: Majalengka, Argamukti, Mt. Ciremai National Park (MZB 13244). 7. Palaina nubigena Möllendorff, 1897, outer Surface of operculum, WeSt JaVa: Cianjur, Kandang Badak (ZMA 68468). 8. Arinia yanseni n. sp., inner Surface of operculum, EaSt JaVa: Rembang, Sumber PacuSen (MZB 18882). Scale bar = 0.07 mm for FigS 3 – 6, 0.1 mm for Fig. 7.

FIGURES 15 – 19. DiStribution of Diplommatinidae in JaVa. 15. Diplommatina baliana Fulton, 1899. 16. Diplommatina diplostoma RenSch, 1931, Diplommatina nevilli (CroSSe, 1879) and Diplommatina canaliculata Möllendorff, 1887. 17. Diplommatina auriculata Möllendorff, 1897. 18. Palaina gedeana Möllendorff, 1897, Palaina nubigena Möllendorff, 1897 and Arinia yanseni n. sp. 19. Opisthostoma javanicum Van Benthem Jutting, 1932, Opisthostoma uranoscopium Van Benthem Jutting, 1932 and Opisthostoma sp.

FIGURES 46 – 48. Diplommatinidae from JaVa. 46. Palaina gedeana Möllendorff, 1897, JaVa (lectotype SMF 105100). 47. Palaina nubigena Möllendorff, 1897, WeSt JaVa: Cikuray (lectotype SMF 105095). 48. Arinia yanseni n. sp., EaSt JaVa: Rembang, Sumber PacuSen (holotype MZB 18893). Scale bar = 1.0 mm.