Hogna ingens (Blackwall, 1857)
Crespo, Luis C., Silva, Isamberto, Enguidanos, Alba, Cardoso, Pedro & Arnedo, Miquel, 2022, Island hoppers: Integrative taxonomic revision of Hogna wolf spiders (Araneae, Lycosidae) endemic to the Madeira islands with description of a new species, ZooKeys 1086, pp. 84-135 : 84
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|Hogna ingens (Blackwall, 1857)|
Lycosa ingens Blackwall, 1857: 284 (Df).
Lycosa ingens Blackwalli, 1867: 203 (Dm).
Trochosa ingens Kulczynski, 1899: 423, pl. 9, fig. 121 (mf).
Geolycosa ingens Roewer, 1955: 241.
Geolycosa ingens Roewer, 1960: 689, fig. 387e (f).
Hogna ingens Wunderlich, 1992: 459, fig. 720b, fig. 724a.
Holotype: no type materials from the Blackwall collection were found neither at the OUMNH nor the NHM.
Deserta Grande • Vale da Castanheira (N), 1 ♀ ( SMF21994 View Materials ), 26.III.1967, 1 ♀ (CRBALC0591) and 4 juveniles (CRBALC0593, CRBALC0594, CRBALC0595, CRBALC0592), 32.56685°N, 16.53694°W, 25.III.2017, hand collecting, leg. L. Crespo; (unknown location), 3 ♀♀ (MNHNP AR16186) GoogleMaps .
Hogna ingens can be diagnosed from all other Madeiran Hogna by the aspect of its legs, blackish with white patches (Figs 13 View Figure 13 , 26C View Figure 26 ), and additionally by its genitalia. In males, according to literature, by the inclined palea shield ( Wunderlich 1992: 596, fig. 720f). In females, by short epigynal anterior pockets, with lateral borders divergent and anteriorly swollen median septum (Fig. 12A View Figure 12 ).
Male: We could not examine any male specimens.
Female (CRBALC0591): (Fig. 12 View Figure 12 ). Total length 25.1; carapace: 14.8 long, 11.0 wide.
Colour: carapace greyish brown, densely covered with short black setae, with a cream longitudinal band present from fovea to posterior margin of carapace; with two faint light grey marginal bands suffused with black patches, covered with white setae; four striae well visible on each flank. Chelicerae black except apically, reddish brown, covered in black setae. Gnathocoxae and labium overall orange-brown, densely covered with black setae; sternum greyish brown, densely covered with black setae. Legs greyish, with a variable number (6-8) of lightly coloured patches covered by white setae. Pedipalps greyish, densely covered in black setae. Abdomen densely covered in black setae, with only four very small white patches dorsally and a small anterolateral band of white setae; venter densely covered in black setae, with only two faint median bands of small white patches.
Eyes: MOQ: MW = 0.7 PW, MW = 1.2 LMP, MW = 1.1 AW; Cl = 0.5 DAME. Anterior eye row slightly procurved.
Legs: Measurements: Leg I: 37.7, TiI: 8.9; Leg IV: 35.9, TiIV: 8.4; TiIL/D: 2.3. Spination of Leg I: FeI: d1.1.0, p0.0.2; TiI: p0.0.0, v2s.2s.2s; MtI: p0.0.1, r0.0.1, v2s.2s.1s. MtI an TiI with dense scopulae.
Epigyne: anterior pockets far apart, short, with lateral borders anteriorly convergent, then becoming divergent (Fig. 12A View Figure 12 ); anterior pocket cavities shallow; median septum anteriorly swollen, with wide posterior transverse part (Fig. 12A View Figure 12 ); spermathecae moderately swollen (Fig. 12B View Figure 12 ); copulatory ducts basally with a laterally projected bulbus (Fig. 12B View Figure 12 ); fertilisation ducts emerging at the base of copulatory duct (Fig. 12B View Figure 12 ).
This species is known only from Vale da Castanheira, a 1 km2 valley in the north end of Deserta Grande (Fig. 14 View Figure 14 ).
Vale da Castanheira is a semi-arid grassland area.
Hogna ingens was declared Critically Endangered in previous works ( Cardoso 2014; Crespo et al. 2014b). Its restricted habitat has been subject to biological invasions since humans set foot in Deserta Grande, with the introduction of herbivore vertebrates and, more recently, of the herb Phalaris aquatica L., which grows abundantly throughout the valley, limiting the access of H. ingens to shelter below rocks and fissures and displacing native flora. A recovery program of the valley’s vegetation is being conducted, and recent data indicates the spider population is increasing. An ex-situ breeding program is currently being conducted by the Bristol Zoo to safekeep populational levels.
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