Dipturus Rafinesque, 1810a

Ebert, David A., White, William T., Ho, Hsuan-Ching, Last, Peter R., Nakaya, Kazuhiro, Séret, Bernard, Straube, Nicolas, Naylor, Gavin J. P. & De Carvalho, Marcelo R., 2013, An annotated checklist of the chondrichthyans of Taiwan, Zootaxa 3752 (1), pp. 279-386 : 352-354

Genus Dipturus Rafinesque, 1810a

Longnosed Skates / Ħḓǎé

Dipturus Rafinesque, 1810a: 16 . Type species: Raja batis Linnaeus, 1758 , by monotypy.

Dipturus gigas ( Ishiyama, 1958b)

Giant Skate / RĦḓǎ

Raja (Tengujei) gigas Ishiyama, 1958b: 386 , figs. 84, 86. Holotype: FUMT P1558 , adult male 1345 mm TL, off Aichi Prefecture, Kumano-nada , Japan.

Local synonymy: New record for Taiwan.

Taiwan voucher material: (3 spec.) CSIRO H 7395-15 (tissue accession GN9998 ), female 299 mm TL, Da-xi, 14 Mar 2012 ; NMMB-P HO-316 , Da-xi, 15 Apr 2012 ; NMMB-P HO-346 , Da-xi, 5 May 2012 .

Remarks: One of us ( DAE) observed this species infrequently at the Da-xi during a survey in May 1988 (identification confirmed by H. Ishihara). Frequently confused with D. macrocauda , particularly at juvenile stages.

Conservation status: Data Deficient.

Dipturus kwangtungensis ( Chu, 1960)

Kwangtung Skate / ǠΦĦḓǎ

Raja kwangtungensis Chu, 1960: 148 , figs. 137–139. Holotype: SFI 57-0676 , juvenile male 336 mm TL, Jia-Bo, East China Sea, China.

Local synonymy: Raja kenojei (not of Muller & Henle): Teng, 1962: 211; Chen & Chung, 1971: 20, fig. 14; Shen, 1984a: 74, fig.; Shen, 1984b: 6, pls 10-1, a, b; Chen & Yu, 1986: 149; Okada et al., 1935:59; Ishiyama, 1958a: 14. Raja (Dipturus) kwangtungensis: Chen & Joung, 1989: 3-4 , fig. 1; Chen & Joung, 1993: 75, pl. 10 (fig. 1). Dipturus kwangtongensis: Shao et al., 2008: 237 . Dipturus kwangtungensis: Shen & Wu, 2011: 96 , fig.

Taiwan voucher material: (23 spec.) CSIRO H 7394-11 , adult male 626 mm TL , CSIRO H 7394-12 (tissue accession GN9928 ), female 753 mm TL , HUMZ 213840 (tissue accession GN9931 ), adult male 600 mm TL , MNHN 2013-0479 (tissue accession GN9929 ), female 610 mm TL , MNHN 2013-0480 (tissue accession GN9930 ), adult male 613 mm TL, Da-xi, 13 Mar 2012 ; CSIRO H 7395-17 , juvenile male 232 mm TL , CSIRO H 7395-18 , female , CSIRO H 7395-19, 240 mm TL, CSIRO H 7395-20 , female 263 mm TL , CSIRO H 7395-21 , juvenile male 301 mm TL , CSIRO H 7395-22 (tissue accession GN9999 ), female 497 mm TL , CSIRO H 7395-23 (tissue accession GN10000 ), juvenile male 419 mm TL , MNHN 2013-0488 (tissue accession GN10001 ), juvenile male 304 mm TL , MNHN 2013-0489 (tissue accession GN10002 ), female 245 mm TL, Da-xi, 14 Mar 2012 ; CSIRO H 7396-04 , juvenile male 180 mm TL, Nan-fang-ao, 14 Mar 2012 ; CSIRO H 7398-05 , female , CSIRO H 7398-06 , juvenile male , CSIRO H 7398-07 , juvenile male, 22°26' N, 120°30' E, Dong-gang, 18 Mar 2012 ; CSIRO H 7401- 07 , juvenile male, 22°26' N, 120°30' E, Dong-gang, 19 Mar 2012 ; CSIRO H 7402-02 , juvenile male 430 mm TL , CSIRO H 7402-04 (3 spec.), 22°26' N, 120°30' E, Dong-gang, 20 Mar 2012 .

Remarks: The literature has continually mixed this species with O. kenojei and O. meerdervoortii . A reexamination of type material and redefinition of these taxa is needed to resolve this nomenclatural confusion.

Conservation status: Data Deficient.

Dipturus macrocauda ( Ishiyama, 1955)

Bigtail Skate / RṚĦḓǎ

Raja macrocauda Ishiyama, 1955: 43 , figs 1–2. Holotype: male 997 mm, Miya fish market, Aichi Prefecture, Japan, 19 Mar 1952; whereabouts not verified.

Local synonymy: Raja (Dipturus) macrocauda: Chen & Joung, 1989:6 , fig. 3. Raja (Dipturus) macrocauda: Chen & Joung, 1993: 75 , pl. 10 (fig. 2); Dipturus macrocauda: Shen & Wu, 2011: 97 , fig.

Taiwan voucher material: (1 specimen) CAS 235421 , Da-xi, 25 May 2005 ; CSIRO H 7397-02 (tissue accession GN10090 ), female 195 mm TL, Da-xi, 15 Mar 2012 .

Remarks: A single specimen was collected at the landing ground of Da-xi by DAE and WTW in May 2005. This species was infrequently observed at the Da-xi by one of us ( DAE) during a survey in April-May 1988. It was typically taken at depths below 300 m.

Conservation status: Data Deficient.

Dipturus tengu ( Jordan & Fowler, 1903)

Acutenose Skate / XṖĦḓǎ

Raja tengu Jordan & Fowler, 1903: 654, fig. 8. Holotype: SU 12912 (specimen bisected), Matsushima Bay , Sndai, Japan.

Local synonymy: Raja tengu: Chen & Chung, 1971: 18 , fig. 13; Shen, 1984a: 74; Yu, 1988: 9. Raja (Dipturus) tengu: Chen & Joung, 1989: 5-6 , fig. 2; Chen & Joung, 1993: 75, pl. 10 (fig. 3). Dipturus tengu: Chen, 2004: 24 ; Shao et al., 2008: 237; Shen & Wu, 2011: 97, fig.

Taiwan voucher material: (9 spec.) CAS 236051 (tissue accession GN10109 ), female 407 mm TL, Dong-gang , Mar 2012 ; AMNH 258306 (tissue accession GN10127 ), female 284 mm TL , MNHN 2013-0482 (tissue accession GN10128 ), female 198 mm TL; NMMB-PHO-242 (tissue accession GN10129 ), adult male 1051 mm TL, 22°26' N, 120°30' E, Dong-gang , 19 Mar 2012 ; CSIRO H 7402-03 (tissue accession GN10151 ), juvenile male 500 mm TL ; NMMB-P HO-265 (tissue accession GN10152 ), female 198 mm TL , MNHN 2013-0483 (tissue accession GN10153 ), juvenile male 210 mm TL, Dong-gang , 20 Mar 2012 ; MNHN 2013-0484 , juvenile male , MNHN 2013- 0485 , female, off Kaohsiung, 320 m depth, 21 Mar 2012 . Drawing in Chen & Chung (1971) is of THUP 02668 (as Raja tengu ), specimen location not verified .

Remarks: Frequently confused with D. gigas . These species need to be further investigated and redescribed.

Conservation status: Data Deficient.

Dipturus wuhanlingi Jeong & Nakabo, 2008

Wuhanling Skate / ùkĦḓǎ

Dipturus wuhanlingi Jeong & Nakabo, 2008: 184 , figs 1-7. Holotype: SFC D 01113 , juvenile male 668 mm TL, southern East China Sea, Apr 1961.

Local synonymy: New record for Taiwan.

Taiwan voucher material: No specimens examined during the present survey.

Remarks: One of us ( DAE) along with H. Ishihara has observed specimens at the Da-xi. Presence in the region needs confirmation as more than one species with this thorn distribution exists in the region.

Conservation status: Data Deficient.