Stomodes tolutarius, : Wagner, 1912

Romαn Borovεc & Pεtεr Hlαváč, 2018, Review of the Stomodes tolutarius species group (Coleoptera: Curculionidae: Entiminae: Otiorhynchini) from the Balkan Peninsula with description of two new species from Greece, Zootaxa 4497 (1), pp. 111-126 : 123

Key to the species of S. tolutarius species group

1. Elytra (Figs 23, 31) with apex not visible in dorsal view, hidden by slope overhanging apex; pronotum with punctures of two different sizes, with unevenly scattered coarse and conspicuously smaller and fine punctures.......................... 2

- Elytra (Fig. 9) with apex visible in dorsal view; pronotum with punctures of equal size, in some species weakly variable in size but never distinctly small............................................................................... 3

2. Rostrum 1.61–1.67× as wide as long; pronotum 1.03–1.06× as wide as long; elytra (Fig. 31) laterally flat, slope overhanging apex, evenly descending downwards; tarsi with segment 2 1.1–1.2× as wide as long.............. S. tolutarius Schoenherr

- Rostrum 1.85× as wide as long; pronotum 1.11× as wide as long; elytra (Fig. 23) with slope overhanging apex, rounded and then perpendicularly falling downwards; tarsi with segment 2 1.4× as wide as long............... S. marocanus Hoffmann

3. Pronotum shiny, with very fine and sparse punctures; distance between two nearest pronotal punctures greater than diameter of a puncture; elytra at most 1.68× as long as wide, with semierect setae on disc, and with finely punctate striae; distance between elytral punctures greater than a puncture diameter; interstriae distinctly wider than striae; funicle segments in females wider than long................................................................................ S. muelleri Lona

- Pronotum matt, densely punctate; distance between two nearest pronotal punctures smaller than diameter of a puncture; elytra at least 1.81× as long as wide, with semi-appressed setae on disc, and with coarsely punctate striae; distance between elytral punctures shorter than their diameter; interstriae weakly wider or equally wide as striae; funicle segments in females longer than wide or as long as wide (except for S. leonhardi )......................................................... 4

4. Epifrons (Fig. 16) wide, with distinctly concave sides, posteriorly significantly enlarged, at base only slightly narrower than space between anterior margins of eyes; funicle segments 3–7 in females wider than long; pronotum in female wider than long.................................................................................... S. leonhardi Wagner

- Epifrons (Fig. 19) slender, at basal half subparallel-sided, at base significantly narrower than space between anterior margins of eyes; funicle segments 3–7 in females isodiametric or longer than wide; pronotum in female isodiametric or longer than wide................................................................................................ 5

5. Femora ventraly with small but distinct tooth ( Fig. 2 ); rostrum with slightly concave sides and slightly laterally projecting pterygiae; striae and interstriae with equally large punctures; interstriae weakly wider than striae; female sternite VIII (Fig. 3) with isodiametric plate, narrowly sclerotised along anterior margin and with large fenestra; spermatheca (Fig. 4) with ramus and nodulus perpendicular................................................................. S. benedikti sp. n.

- Femora adentate (Fig. 8); rostrum with straight sides, without laterally projecting pterygiae; punctures in striae distinctly larger than those on interstriae; striae and interstriae equally wide; plate of female sternite VIII (Figs 10, 20) wider than long, with apical half uniformly sclerotized and lacking fenestra; spermatheca (Fig. 12) with ramus and nodulus divergent, V-shaped.. 6

6. Epifrons with longitudinal median stria; pronotum widest slightly before the middle; pronotal disc with well isolated punctures; elytra longer, 1.89–1.98× as long as wide; rostrum longer, 1.55–1.67× as wide as long; metatarsal segment 2 1.1× as wide as long; penis (Fig. 13) in dorsal view at apex evenly regularly tapered, tip shortly subtruncate; plate of female sternite VIII (Fig. 10) with apical margin concave................................................... S. dodocunevi sp. n.

- Epifrons convex, lacking stria; pronotum widest at middle; pronotal disc with space between punctures very narrow; elytra shorter, 1.81–1.91× as long as wide; rostrum shorter, 1.78–1.91× as wide as long; metatarsal segment 2 1.4× as wide as long; penis in dorsal view (Fig. 21) with apex weakly enlarged and then tapered and pointed apicad; plate of female sternite VIII (Fig. 20) with round apical margin........................................................... S. letzneri Reitter

Figures

FIGURES 1–2. 1—Stomodes benedikti, habitus; 2—Stomodes dodocunevi, habitus.