Pterostichus malkini

Labonte, James R., 2013, Pterostichus (Anilloferonia) diana LaBonte (Coleoptera: Carabidae: Pterostichini), a replacement name for P. (A.) lanei (Hatch, 1935), and validity and redescription of P. (A.) malkini (Hatch, 1953), Zootaxa 3682 (4), pp. 563-571 : 566-568

Distinguishing Pterostichus malkini from other species of P. ( Anilloferonia )

Four characters readily distinguish P. malkini from P. diana and P. testacea : the number of supraorbital setae, the shape of the male metafemur, the form of the last visible abdominal ventrite of the male just anterior of the apical margin, and the form of the median lobe of the aedeagus. There are other distinguishing characters but these are subtle, are often variable, and sometimes cannot be accurately assessed without access to reliably identified reference specimens. They will not be addressed herein as that would be more appropriate in a broader treatment of Anilloferonia .

All known specimens (6) of P. malkini have only one pair of supraorbital setae ( Figure 2 ). All specimens of P. diana (111) ( Figure 3 ) and P. t e s t a c e a (21) ( Figure 4 ) I examined have two pairs of supraorbital setae, with the exception of one specimen of P. diana with two setae on one side and three on the other.

The median posterior margins of the metafemora of the three male P. malkini examined are obliquely angulate and the ventral faces of the metafemora are broader at that point than at either end ( Figure 5 ). The posterior margins of the metafemora of female P. malkini ( Figure 6 ) and both females and males of P. diana (male, Figure 7 ) and P. testacea (male, Figure 8 ) are shallowly, smoothly arcuate throughout and the metafemora are more or less narrowly cylindrical throughout.

The last abdominal ventrite in the three available specimens of male P. malkini has a small, ventrally projecting ridge between the pair of paramedial preapical setae ( Figures 9–10 ). Female P. malkini lack this ridge ( Figure 11 ), although a small preapical protuberance is present. The last abdominal ventrites of females and males of P. diana (male, Figure 12 ) and P. testacea (male, Figure 13 ) lack preapical ridges or protuberances.

The shape of the median lobe of the aedeagus of P. malkini differs from those of P. diana and P. t e s t a c e a as follows: the median lobe is shorter and less sinuate in lateral view ( Figures 14–19 ) and dorsal view ( Figures 20– 22 ), the right ventrolateral tumidity is much smaller and is very close to the base of the median lobe (versus large and more medially located) ( Figures 17–19 ), the meeting of the shaft of the median lobe and the base is narrowly perpendicular in right lateral perspective ( Figure 17 ) versus being arcuate and broader ( Figures 18–19 ), the apex is shorter in both lateral and dorsal perspectives ( Figures 14–22 ) and in dorsal perspective, the apex is less twisted to the anatomical left and the tip of the apex is more rounded ( Figures 23–25 ).

Figures

FIGURES 2 – 4. Supraorbital setae of species of Pterostichus (Anilloferonia). Figure 2, P. malkini. Figure 3, P. d i a n a. Figure 4, P. t e s t a c e a.

FIGURES 5 – 8. Metafemora of species of Pterostichus (Anilloferonia). Figure 5, male P. malkini. Figure 6, female P. malkini. Figure 7, male P. d i a n a. Figure 8, male P. testacea.

FIGURES 9 – 13. Last abdominal ventrite of species of Pterostichus (Anilloferonia). Figure 9, lateral view, male P. malkini. Figure 10, ventral view, male P. malkini. Figure 11, female P. malkini. Figure 12, male P. d i a n a. Figure 13, male P. testacea.

FIGURES 14 – 16. Left lateral views of median lobe of aedeagus of species of Pterostichus (Anilloferonia). Figure 14, P. malkini. Figure 15, P. d i a n a. Figure 16, P. t e s t a c e a.

FIGURES 17 – 19. Right lateral views of median lobe of aedeagus of species of Pterostichus (Anilloferonia). Figure 17, P. malkini. Figure 18, P. d i a n a. Figure 19, P. t e s t a c e a.

FIGURES 20 – 22. Dorsal views of median lobe of aedeagus of species of Pterostichus (Anilloferonia). Figure 20, P. malkini. Figure 21, P. d i a n a. Figure 22, P. testacea.

FIGURES 23 - 25. Dorsal views of apex of median lobe of aedeagus of species of Pterostichus (Anilloferonia). Figure 23, P. malkini. Figure 24, P. d i a n a. Figure 25, P. t e s t a c e a.