Pterostichus (Anilloferonia) malkini (Hatch)

Labonte, James R., 2013, Pterostichus (Anilloferonia) diana LaBonte (Coleoptera: Carabidae: Pterostichini), a replacement name for P. (A.) lanei (Hatch, 1935), and validity and redescription of P. (A.) malkini (Hatch, 1953), Zootaxa 3682 (4), pp. 563-571 : 564-566

Redescription of Pterostichus (Anilloferonia) malkini (Hatch)

( Fig.1 )

Color: Rufous to dark rufous throughout. Total body length (mandible apices to occiput + middle of pronotum from anterior margin to posterior margin + base of scutellum along suture to elytral apex): 6.2–7.8 mm. Head: Mandibles elongate, with sharp, hooked apices. Frontolateral carinae distinct, well-defined, parallel except anterior of antennal insertions, where each forms an arcuate obtuse angle with the medially directed anteriormost portion; each extending from frontoclypeal suture to supraorbital seta and highest at antennal insertion. Each frontal impression distinct and deeply impressed, extending from posterior half of clypeus about halfway to occiput, straight on posterior half of clypeus and then arcing from antennal insertion mediad, becoming arcuately divergent at posterior extent. Moderately wide, at widest point 0.6–0.7 as wide as widest point of pronotum. Antennae extending about two segments beyond posterior margin of pronotum. Eyes minute, 0.07–0.125 mm long at greatest length, narrowly ovate and from 0.05–0.075 mm wide, facets faintly visible. One pair of supraborbital setae, anterior pair absent. Pronotum: Greatest width 1.1 times greater than length along midline, greatest width about 0.7 of total length from posterior margin, just posterior of anterolateral setae. Anterior angles distinctly protruding beyond truncate anteromedial margin, narrowly rounded at approximate acute angle. Anterolateral setae about 0.75 of total length from posterior margin and adjacent to lateral margin. Lateral margins evenly slightly convex from anterior angles to about midlength of pronotum, thereafter straight and slightly convergent to posterior angles (varying to faintly sinuate at about anterior limits of posterlateral depresssions and parallel just anterior of posterior angles). Posterior angles about 90 0 or narrowly obtuse, narrowly rounded. Posterolateral setae in posterior angles, about twice distance from lateral margins as posterior margin. Posterior margin faintly convex laterad of each posterolateral impression, faintly emarginotruncate between impressions. Lateral explanations more or less evenly narrow from anterior angles to anterolateral setae, then gradually broadening to about anterior limits of posterolateral impressions, rapidly broadening thereafter until reaching posterior margin. Anteromedial impression at most very faint, indicated by row of shallow, large punctures. Median line strongly impressed, extending from about anteromedial impression to just anterior of or reaching posterior margin. Dorsal surface shallowly evenly convex either side of median line to about 0.6 distance to lateral margin, somewhat flattened laterad from there. Posterolateral impressions linear, deeply incised. Dorsal surface between median line and impressions somewhat flattened, but tumid compared to flat or faintly concave areas laterad of impressions. Impressions extend 0.3–0.4 total pronotal length from posterior margin. Areas between median line and impressions with sparsely to moderately dense fine and coarse deep punctures, areas laterad of impressions with sparse fine shallow punctures, sometimes virtually impunctate. Posterior margin with complete marginal bead. Elytra: Total length about 1.7 times combined greatest width of elytra. Anterior margins slightly concave. Humeral teeth from very small and barely visible to sharp and distinct. Expression of parascutellar striae and angular bases of striae 1 highly variable among specimens, including asymmetrical expression in some individuals. Parascutellar stria may coalesce with stria 1, be distinct but not coalescent with stria 1, or be intermittent and not coalescent with stria 1. Angular base of stria 1 may coalesce with stria 1 or be discontinuous with the anterior portion of stria 1. No basal punctures or setae are present. Striae with faint, evenly spaced, elongate punctures. Epipleuron with distinct, narrow, preapical plica, not denticulate at plica. Umbiculate series of stria 8 with five setiferous punctures in anterior series, one at about anterior third, a pair behind middle, a triad anterior of plica. A pair of subapical umbiculate setiferous punctures in stria 7 posterior of plica. Ventral thoracic sclerites: Prosternal process varying from faintly to strongly margined. Prosternum impunctate except for variable scattered coarse, deep punctures along lateral portions. Proepisterna with moderately dense coarse, deep punctures concentrated in anterior 2/3. Mesosternum impunctate. Mesoepisterna with dense coarse, deep punctures, most in anterior half along mesepisternal depression. Metasternum impunctate except along lateral portions, which have moderately dense coarse, deep punctures. Metaepisterna with variably distributed scattered coarse, deep punctures. Abdominal ventrites: Ventrites 1 and 4– 6 impunctate. Ventrites 2 and 3 with variable scattered punctures about midway between median and lateral margins. Ventrites 3–5 with a pair of paramedial setae. Female ventrite 6 with two pairs of apical paramedial setae. Male ventrite 6 with a single pair of apical paramedial setae. Male ventrite 6 with a low, vertically directed lobe between the setae and just anterior of posterior margin, this lobe broadly truncate when viewed from purely anterior or posterior perspectives. Female ventrite 6 without a lobe, but with a very small tumidity between the median pair of paramedial setae. Legs: Female protarsomeres 2–4 broadly triangular and lacking ventral squamose setae, protarsomeres 1 and 5 moderately elongate, 5 glabrous ventrally. Male protarsomeres 1 moderately elongate and expanded in apical half, 2–4 larger and broader than in female, 5 as in female; ventral squamose setae on expanded portion of 1 and 2–3, 4 and 5 lacking such setae. Mesotarsomeres 2–4 in both females and males somewhat short and broad, rest moderately elongate. Metatarsomeres in both females and males moderately elongate. Metatrochanters slender and cylindrical, narrowly rounded at apex. Female metafemora slender, more or less cylindrical, ventral margin smooth and shallowly, evenly convex. Female metatibiae straight in dorsal view.

Male metafemora flattened and broadest in anterior and posterior views, with distinct obtuse angle at about 2/3 along ventral margin from femoral base, femoral base about 1/4 as wide and apex about half as wide as at angle; ventral face with very well defined carinae on either side extending from angle to apex and delimiting a concavity, carinae merging medially at angle to form interior delimitation of concavity. Male metatibiae slightly arcuate in dorsal view. Aedeagus: In lateral view, ventral margin of median lobe meets base at right angle. Right medioventral tumidity of median lobe very small, near base, thus in lateral view ventral margin of median lobe unisinuate with a single concavity near apical third. In dorsal view, tip of median lobe very short, rounded at apex, strongly convex at right extreme and straight along entire apical half of median lobe. Internal sac with a single large, slender sclerotized spine.

Figures

FIGURE 1. Habitus of male Pterostichus (Anilloferonia) malkini (Hatch).