Diaparsis (Diaparsis) demeter Khalaim, 2022

Khalaim, A. I., 2022, Two new species of Tersilochinae from Madagascar (Hymenoptera: Ichneumonidae: Tersilochinae), Russian Entomological Journal 31 (2), pp. 163-167 : 164

Diaparsis (Diaparsis) demeter Khalaim , sp.n.

MATERIAL EXAMINED. Holotype: ♀ ( MNHN), Madagascar, Anamalanga Reg., Rogez [Andekaleka], Forêt cote Est, XII.1932, coll. A. Seyrig, “ Telcocrypta telescopica Srg. TYPE”.

Paratypes. Madagascar, Anamalanga Reg., Rogez [Andekaleka], Forêt cote Est, 1935, coll. A. Seyrig. Same locality and collector, X.1936, 1 ♀ ( ZISP) . Same locality and collector, X.1936, 1 female ( MNHN) . Perinet, Forêt cote Est, III.1932, coll. A. Seyrig, “ Telcocrypta telescopica perinetisana Srg. TYPE”, 1 female ( MNHN) .? Betsiboka Reg., “An ... [hand-written illegible text]” (Andriba? [Andrioa]), XI.1933, coll. A. Seyrig, 1 ♀ ( MNHN) .

Non-type material. Madagascar, Anamalanga Reg., Rogez [Andekaleka], Forêt cote Est, XII.1932, coll. A.Seyrig, 1 ♀ ( MNHN) .

COMPARISON. Diaparsis demeter sp.n. is very similar to D. heterocoloides sp.n. but differs from this species by the colour pattern of mesosoma, distinct notaulus, deep and broad foveate groove of mesopleuron with distinct transverse wrinkles reaching the epicnemial carina anteriorly, and mandible with upper tooth almost 2.5 times longer than the basal mandibular width.

DESCRIPTION. Female. Body length 10.5 mm. Fore wing length about 5.7 mm.

Head with gena roundly constricted posterior to eyes; gena in dorsal view about 0.65 times as long as eye width. Clypeus large, lenticular, smooth, with fine or distinct punctures in upper 0.7, separated from face by weak and broad impression. Mandible moderately slender, weakly constrict- ed in basal half, with lower tooth about 2.5 times as long as the upper tooth. Malar space 0.4–1.0 times (almost 1.0 times in holotype) as long as basal mandibular width. Flagellum in all specimens with complete antennae with 27 flagellomeres, distinctly tapered towards apex; subbasal flagellomeres 1.3– 1.6 times as long as broad, subapical flagellomeres 1.3 times as long as broad (in holotype) to square; flagellomeres 2 or 3 to 7 bearing subapical finger-shaped structures on outer surface. Face with weak median swelling. Face and frons with dense punctures, smooth or very shallowly granulate between punctures. Vertex and gena with fine and moderately dense punctures; vertex subpolished and gena polished between punctures. Occipital carina complete, evenly convex mediodorsally.

Mesoscutum densely punctate, shallowly granulate or polished between punctures. Notaulus weakly impressed, with distinct longitudinal wrinkle or irregular wrinkles on anterolateral side. Scutellum with lateral longitudinal carinae present at its anterior 0.2–0.3. Upper end of epicnemial carina abruptly curved to reach front margin of mesopleuron at level of lower 0.3 of pronotum. Foveate groove of mesopleuron moderately deep, broad, oblique, situated in anterior 0.6 of mesopleuron, with distinct transverse wrinkles usually reaching epicnemial carina anteriorly. Mesopleuron densely punctate on smooth or weakly granulate background, with impunctate area in centre (above foveate groove). Propodeal spiracle separated from pleural carina by 1.0–2.0 times diameter of spiracle. Propodeum with dorsolateral areas punctate on smooth or weakly granulate background; basal keel 0.2–0.3 times as long as apical area. Apical area flat or weakly impressed along midline, more or less pointed anteriorly, uneven, weakly shining; apical longitudinal carinae strong, complete, reaching transverse carina anteriorly.

Fore wing with second recurrent vein (2m-cu) postfurcal. First abscissa of radius (Rs+2r) straight, much longer than width of pterostigma. First and second sections of radius (Rs+2r and Rs) meeting at slightly acute angle. Intercubitus (2rs-m) slightly thickened, longer than abscissa of cubitus between intercubitus and second recurrent vein (abscissa of M between 2rs-m and 2m-cu). Metacarpus (R1) almost reaching tip of the wing. Hind wing with nervellus (cu1&cu-a) weakly reclivous. Legs slender, tarsal claws not pectinate.

Metasoma unusually long, posterior to second tergite strongly compressed laterally. First tergite very slender, about 6.2 times as long as posteriorly broad, trapeziform to almost round in central cross-section, smooth, sometimes with fine longitudinal striae laterally before glymma; glymma small (sometimes indistinct), situated slightly behind centre of tergite; in dorsal view, petiole with lateral margins subparallel, postpetiole widened anteriorly and distinctly broader than petiole; upper margin of first tergite, in lateral view, nearly straight in anterior 0.7 and weakly arcuate in posterior 0.3. Second tergite 4.0–5.0 times (almost 5.0 times in holotype) as long as anteriorly broad. Thyridial depression very long, over 5.0 times as long as broad, with posterior end rounded or pointed. Ovipositor slender and exceptionally long (longer than body), evenly bent upwards, with a slight dorsal subapical swelling.

Head black; clypeus yellowish brown in lower 0.4–0.5 and dark reddish black in upper part to almost entirely reddish brown. Mouthparts and mandibles (teeth dark red) brownish yellow to yellow-brown. Antenna entirely dark brown to brownish black; in holotype flagellum yellow-brown, probably due to discolouration. Mesosoma reddish brown with upper part of pronotum darkened, and mesoscutum, scutellum and postscutellum black; propodeum sometimes narrowly darkened anteriorly. Tegula yellow-brown or brown. Pterostigma brown. All legs rather uniformly brownish yellow; hind leg sometimes slightly darkened with brown. Metasoma entirely pale brown to brown, sometimes first tergite entirely or only its postpetiole dark brown.

Male. Unknown.

VARIATION. One non-type female differs from typical specimens by its retatively short ovipositor which is about as long as the body, mandible with rounded ridge along its lower margin centrally, upper tooth of mandible much broader and almost 3.0 times longer than the lower tooth, mesopleuron with narrow and sharp foveate groove in its centre (foveate groove does not reach the epicnemial carina anteriorly), and mesosoma less black marked (mesoscutum with only black mark on anterior half of central lobe; scutellum, postscutellum and front margin of propodeum weakly darkened).

ETYMOLOGY. The species name derives from Demeter (noun), the Olympian goddess of the harvest and agriculture in ancient Greek mythology.

DISTRIBUTION. Madagascar.