Ulomorpha polytricha Alexander, 1930

Kato, Daichi, Watanabe, Kozo & Kolcsar, Levente-Peter, 2020, Synopsis of the genus Ulomorpha Osten Sacken, 1869 (Diptera, Limoniidae) in Japan, ZooKeys 999, pp. 147-163 : 147

Ulomorpha polytricha Alexander, 1930 Figs 4 , 5A

Ulomorpha polytricha Alexander 1930a: 72 (type locality: Japan, Nansei Islands, Yaku-shima I., Kosugidani); Alexander 1930b: 508; Nakamura 2014: 20; Oosterbroek 2020.

Material examined.

Holotype. ♂, Japan, Nansei Islands, Yaku-shima I., Kosugidani; altitude 2,500 feet; 30°20.84'N, 130°35.25'E (rough coordinate); 29 Apr. 1929; S. Issiki leg. (USNM).

Non-types. Japan: [Nansei Islands] Yakushima I.: • 17♂, 1♀; near Shirataniunsui-kyô Valley; alt. 600 m; 30°23.04'N, 130°34.37'E; 25 Apr. 2019; D. Kato leg. (BLKU) • 1♂; same data as previous except 27 Apr. 2019 • 1♂; unnamed stream near Kigensugi; alt. 1270 m; 30°18.1'N, 130°32.56'E; 25 Apr. 2019; D. Kato leg. (BLKU) • 2♂, 1♀; unnamed stream near Mt Mae-dake; alt. 625 m; 30°18.91'N, 130°36.61'E; 25 Apr. 2019; D. Kato leg. (BLKU).

Diagnosis.

Body blackish. Vertex sparsely pruinose. Basal 2-3 flagellomeres oval, with pubescence ventrally. Prescutum sparsely pruinose. Wing brownish tinged, unpatterned except indistinctly darker stigma. Halter yellow. Interbase with outer lobe shorter than medial lobe in dorsal view, medial lobe medial to base of outer lobe more than twice as long as wide. Aedeagus with rod-shaped distal part, about 2.2 times as long as wide and almost straight.

Description.

Male. Body length 5.7-8.0 mm, wing length 6.0-8.1 mm.

Head: subnitidous black, sparsely dusted with gray pruinosity; vertex with brighter gray pruinosity at anterior end. Rostrum and mouthparts dark brown. Antenna brown; scape and pedicel sometimes slightly darker; basal 2-3 flagellomeres oval, ventrally with pubescence (Fig. 4B ).

Thorax: subnitidous dark brown to black, sparsely dusted with brownish pruinosity; postpronotum yellowish or brownish. Wing (Fig. 4C ) tinged with brown;, basal region yellowish; stigma indistinctly darker; veins dark brown, yellowish on basal part of wing; barely dark seam sometimes present on fork of Rs to crossvein r-m. Halter yellow. Legs mainly yellow to dusky yellow; fore coxa dark on basal half; femora narrowly dark at tips of mid and hind pairs; fore femur dark on distal 2/3; tibiae narrowly dark at tips; tarsi weakly dark from tip of tarsomere 1 to apical segment.

Abdomen: subnitidous dark brown to black, sparsely covered with brownish pruinosity.

Male terminalia (Fig. 4D-H ): caudal margin of tergite 9 roundly produced at middle, with shallow U-shaped notch at center. Outer gonostylus in dorsal view with tip narrowed and strongly curved anteriorly, concaved margin with small teeth (Fig. 4E ). Interbase with outer lobe shorter than medial lobe in dorsal view (Fig. 4F ); outer lobe weakly wide at base, inner basal end at basal 1/3 of medial lobe (Fig. 4H ); medial lobe gradually narrowed distally, distal part medial to base of outer lobe more than twice as long as wide (Fig. 4H ). Aedeagus with rod-shaped distal part, about 2.2 times as long as wide and almost straight (Fig. 4G ).

Female. Body length 7.2-7.6 mm, wing length 6.7-7.1 mm. Generally resembling male.

Ovipositor: dark brown; yellow on cercus, hypogynial valve, and distal 1/4 of tergite 10; cercus twice as long as tergite 10.

Distribution.

Japan (Nansei Islands: Yakushima Island) (Fig. 5A ).

Remarks.

This species is similar to U. amamiana sp. nov. See the key to the Japanese species above for differentiation and diagnostic characters.

Figures

Figure 4. Ulomorpha polytricha Alexander, 1930. A habitus, male B basal part of antenna, lateral view C wing D male genitalia, dorsal view E tip of outer gonostylus, dorsal surface F aedeagal complex, dorsal view G aedeagal complex, lateral view H interbase, posterodorsal view. Scale bars: 3 mm (A, C); 0.1 mm (B, D-H).

Figure 5. Distribution map of Japanese species of Ulomorpha. A U. amamiana Kato & Kolcsar, sp. nov. (orange), U. nigricolor Alexander, 1924 (blue), U. polytricha Alexander, 1930 (green) B U. longipenis Kato & Kolcsar, sp. nov. (red).