Harperocallis robustior (Steyerm.) L.M. Campb. & Dorr, 2013

Campbell, Lisa M. & Dorr, Laurence J., 2013, A synopsis of Harperocallis (Tofieldiaceae, Alismatales) with ten new combinations, PhytoKeys 21, pp. 37-52 : 44-45

Harperocallis robustior (Steyerm.) L.M. Campb. & Dorr comb. nov. Figs 2C, D , 3

Tofieldia sessiliflora var. robustior Steyerm., Fieldiana, Bot. 28 (1): 157. 1951. Type: Venezuela. Lara:Wet meadow at Las Sabanetas, above Los Aposentos, west of Humocaro Bajo, 2530 m, 5 Feb 1944, J.A. Steyermark 55291 (holotype: F [F0046066F!]; isotype: NY!).

Isidrogalvia robustior (Steyerm.) Cruden, Syst. Bot. 16 (2): 278. 1991. Type: Based on Tofieldia sessiliflora var. robustior Steyerm.

Type.

Based on Tofieldia sessiliflora var. robustior Steyerm.

Distribution

( Fig. 1 ). Andes of Venezuela (Lara, Mérida, and Trujillo states); 2200-3700 m.

Representative specimens.

Venezuela. Lara: Mpio. Moran, below Páramo Las Rosas, 2285-3290 m, 25 Jun 1979, R. Liesner et al. 7965 (MO, NY). Mérida: Mpio. Rangel, Cerro el Guamo, 3100-3200 m, 19 Jun 1988, L.J. Dorr & L.C. Barnett 5627 (NY). Trujillo: Páramo de Cachaco, 3400-3700 m, 7 Nov 1927, P. Christ 84 (NY); Mpio. Boconó, Páramo de Guaramacal, ca. 3080 m, ca. 9°14'N, 70°11'W, 28 Apr 1988, L.J. Dorr et al. 4967 (NY, PORT); Páramo de Guirigay, 3000 m, 22 Apr 2002, L.J. Dorr et al. 9200 (NY, PORT, US); Parque Nacional Guaramacal, vertiente norte, ca. 1977-2350 m, 28 Apr 1998, B. Stergios et al. 17327 (PORT-unicate); Laguna Larga via Laguna Las Parias to Laguna Eco, Páramo de Motumbo, 2400-2600 m, 15 Sep 2003, B. Stergios et al. 20368 (F, MO, NY, PORT, US).

Figures

Figure 1. Distribution of Harperocallis.

Figure 2. Habit and habitats of Harperocallis. Harperocallis flava. A Flower (note the peduncular bract, pronounced connective, and tuberculate ovary) B Habitat. Harperocallis robustior. C Inflorescence D Habit and habitat. Harperocallis schomburgkiana. E Flower F Leaves. (A, unvouchered, photograph A. R. Schotz; B, photograph G. Anglin, C, D B. Stergios et al. 20368 (PORT), photograph K. J. Wurdack; E, F K. J. Wurdack et al. 5636 (US), photograph K. J. Wurdack).

Figure 3. Harperocallis robustior. A Habit B Leaf (detail of parallel veins and tomentose margins) C Inflorescence (detail showing peduncular and calycular bracts subtending flowers) D Peduncular bracts (detail) E Flower F Tepal (adaxial view) G Ovary (note the tuberculae) H Ovary (cross-section showing axile placentation and ovules) I Capsule (carpels separating distally). (A, I from L. J. Dorr et al. 4967 (US); B-H from B. Stergios et al. 17327 (PORT).