Fluminicola modoci Hannibal, 1912

Liu, Hsiu-Ping & Hershler, Robert, 2019, A new species and range extensions for three other species of pebblesnails (Lithoglyphidae, Fluminicola) from the upper Klamath basin, California-Oregon, ZooKeys 812, pp. 47-67 : 56

Fluminicola modoci Hannibal, 1912 Figures 4, 8B

Fluminicola modoci - Hannibal 1912: 187, pl. 8: fig. 30 (in part; Fletchers Spring, south end, Goose Lake, California).

Distribution.

Several springs in the Goose Lake basin, California–Oregon ( Hershler and Frest 1996; Hershler 1999; Hershler et al. 2007).

Referred material.

OREGON. Klamath County. USNM 1190095, USNM 1207964, Wood River, south spring source (42.7372N, 121.9775W), USNM 1144333, USNM 1190094, Tecumseh Spring (42.6424N, 121.9432W), USNM 874187, USNM 874935, USNM 1144336, USNM 1144337, USNM 1144942, USNM 1190086, Barkley Spring (42.3822N, 121.8111W), USNM 1190102, Klamath River, east of Boyle Power Station (42.0934N, 122.0964W), USNM 1190087, Ouxy Spring (42.3989N, 121.8235W), USNM 1144390, USNM 1144923, USNM 1190100, Brown Spring (42.4951N, 121.2956W), USNM 1190093, Spring, Klamath Fish Hatchery (42.6519N, 121.9479W), USNM 1190096, Reservation Spring (42.7023N, 121.9629W), USNM 1144407, USNM 1190098, Spring, Williamson River campground (42.6584N, 121.8499W), USNM 1190097, Spring Creek, headspring (42.6690N, 121.8860W), USNM 1144411, USNM 1190090, Crystal Spring (42.5736N, 122.0823W), USNM 1144966, USNM 1190121, Big Springs at Bonanza (42.1982N, 121.4004W), USNM 1144418, USNM 1185800, Casebeer Spring, ouflow at Gerber Dam Road (42.2056N, 121.0592W), USNM 1144454, USNM 1190123, Duncan Spring, north complex (42.0416N, 121.0689W), USNM 1190123, Lost River at Big Springs City Park, Bonanza (42.1976N, 121.4002W), USNM 114469, USNM 1190124, Gwinn Spring Creek at Gwinn Spring Creek Road (42.0063N, 120.9545W), USNM 1020714, USNM 1144673, USNM 1144925, USNM 1154376, Link River under US97/OR140 bridge (42.2185N, 121.7892W). USNM 1190137, Link River, above Link River dam (42.2341N, 121.8036W), USNM 1144494, USNM 1144986, USNM 1207967, Lost River, Stevenson County Park (42.1831N, 121.5994W), USNM 1190139, Lost River, below Harpold Dam (42.1702N, 121.4530W), USNM 1144503, USNM 1190101, Sprague River at Beatty Gap (42.4476N, 121.2377W), USNM 1144515, USNM 1190099, Williamson River, Klamath County park (42.5705N, 121.8791W), USNM 1144520, USNM 1190140, Upper Klamath Lake, south of Modoc Point (42.4373N, 121.8672W). Lake County: USNM 1144565, USNM 1185798, Spring southeast of Slash Spring, Yocum Valley (42.0174N, 120.7316W).

Remarks.

The UKL F. modoci range from 2.6-8.4 mm in shell height and include specimens with eroded spires that closely resemble the type material for this species (e.g., Fig. 4A) as well as individuals with fully intact, ovate-conic to trochoidal shells. Radula: central teeth with 2-6 lateral cusps, 1-4 basal cusps; lateral teeth with 3-7 cusps on outer side, 2-5 cusps on inner side; inner marginal teeth with 13-30 cusps; outer marginal teeth with 20-43 cusps (USNM 1144337, USNM 1144418, USNM 1144925).

The new records extend the range of F. modoci about 130 km westward from the northwestern portion of the Goose Lake basin.

Twenty-seven (27) COI and 31 cytB haplotypes were detected in the UKL specimens of F. modoci (Suppl. materials 2, 3, respectively).

The "Klamath pebblesnail" (also referred to as Fluminicola n. sp. 1) that was recognized by Frest and Johannes in their UKL contract reports (also see Frest and Johannes 1999) and subsequently included in the Northwest Forest Plan as a Survey and Manage species, may correspond to F. modoci .