Brasenia victoria (Casp. 1856) Weberbauer (1893: 374)

Winterscheid, Heinrich, 2020, Typifications in fossil-species of Brasenia, Magnolia, Vitis, and Symplocos from the central European Neogene, Phytotaxa 428 (2), pp. 113-121 : 113-115

Brasenia victoria (Casp. 1856) Weberbauer (1893: 374)

Fig. 2A–D . Basionym:— Holopleura victoria Caspary (1856: 216–217 ; pl. 12, figs 10–22), Caspary in Ludwig (1857: 94–97; pl. 17, figs 6–18). Lectotype (designated here):—[MB.Pb.2017/1474.1!], repository: Museum für Naturkunde, Leibniz-Institut für Evolutions- und

Biodiversitätsforschung, Berlin (MfN). Locus typicus: Germany. Hesse: Wetterau, Bad Salzhausen near Nidda. Stratum typicum:

Middle Miocene (Langhian), intravolcanic brown-coal deposits (“Carpolithen-Kohle von Salzhausen”). Syntypes:—[MB.Pb.2017/1474!] as Holopleura victoria .

Note:—Type not yet cited. The author selects one specimen of the collection no. MB.Pb.2017/1474 (with multiple specimens) as the lectotype.

Occurrence and habitat:— Brasenia victoria occurred in Europe from the late Oligocene to the late Pliocene, in brown-coal deposits and silting up sequences, which reflect fluviatile back-swamp, lake and oxbow lake habitats.

Additional specimens examined:— Germany. Hesse: Wetterau, Bad Salzhausen near Nidda; middle Miocene (Langhian), intravolcanic brown-coal deposits (“Carpolithen-Kohle von Salzhausen”): MfN [ MB.Pb.2017/1482] as Carpolithes oviformis , [ MB.Pb.2017/1483]. MfN, Ludwig coll. [ MB.Pb.1980/0260!]. MfN, P. G. L. A. coll. [ MB. Pb.2005/0615 (orig.-no. 169)!], illustrated in: Kirchheimer (1936: pl. 7, fig. 4a).

= Nymphaea doliolum Ludwig (1860: 123–124; pl. 60, fig. 12a–f), syn. nov. Lectotype (designated here):—[ MB.Pb.2017/1473.1!], repository: Museum für Naturkunde, Leibniz-Institut für Evolutions- und

Biodiversitätsforschung, Berlin ( MfN), Rudolph Ludwig collection. Locus typicus: Germany. Hesse: Wetterau, Bad Salzhausen near

Nidda. Stratum typicum: Middle Miocene (Langhian), intravolcanic brown-coal deposits (“Carpolithen-Kohle von Salzhausen”). Syntypes:—[ MB.Pb.2017/1473!], [ MB.Pb.2017/1479!], [ MB.Pb.2017/1480!], [ MB.Pb.2017/1481!].

Family Magnoliaceae Jussieu (1789: 280) , nom. cons.

Genus Magnolia Linnaeus (1753: 535)

Figures

FIGURE 2. Brasenia victoria (Casp.) Weberb.,Salzhausen near Nidda (Germany); middleMiocene (Langhian) brown coal of intravolcanic sediment deposits. A. Overview of the SEM-stub [no. 2017/1473–1474] with seeds of Brasenia victoria (Casp.) Weberb. (a–c) and Nymphaea doliolum R.Ludw. (d–e): a. Lectotype of Brasenia victoria [MB.Pb.2017/1474.1], e. Lectotype of Nymphaea doliolum [MB. Pb.2017/1473.1]. B. Higher magnification of the Lectotype of Brasenia victoria [MB.Pb.2017/1474.1] with apical cap (embryotheca). C. Brocken open seed (syntype) of Brasenia victoria [MB.Pb.2017/1474.2] with the visible interior multilayered parenchyma (inner testa) and the external columnar sclerotesta. D. Closer view of figure C of the thick-walled columnar sclerotesta.