Cloeodes superior Kluge 1991

Kluge, Nikita J., 2017, Contribution to the knowledge of Cloeodes Traver 1938 (Ephemeroptera, Baetidae), Zootaxa 4319 (1), pp. 91-127 : 123-125

Cloeodes superior Kluge 1991

( Figs 137–141, 143–144 )

Cloeodes superior Kluge 1991: 130 (larva, ♂ and ♀ imagoes, eggs).

Material examined. Holotype: L-S-I ♂ {specimen [III](7)1989}: CUBA, Rio Guama near La-Alcarraza (border of provinces Santiago-de-Cuba and Granma, 60 km W Santiago-de-Cuba), 20°03'N, 76°28'W, 4.III.1989 , coll. N. Kluge. Paratypes: the same locality, 2–8.II.1989: 3 L-S-I ♂, 10 L-S-I ♀, 20 L-S ♂, 10 L-S ♀, 1 larva; Arroyo Paco (tributary of Rio Palma Mocha, 100 km W Santiago-de-Cuba, Prov. Santiago-de-Cuba ), 20°00'N, 76°51'W, 18– 24,1989 , coll. N. Kluge: 3 L-S-I♀, 3 L-S♂, 2 L-S♀, 2 larvae; Titin, 4.XII.1983, coll. C. Naranjo: 20 larvae; Idalia, 25.I.1986, coll. C. Naranjo: 16 larvae.

Additions and corrections to description. Larva. Prostheca of left mandible with 3 blunt processes and 3–4 pointed processes ( Figs 139, 141 ). Maxillary canines and distal dentiseta stout; distal dentiseta widened, with apex somewhat hooked toward canines (as in Figs 8 , 142 ). Other characters as described by Kluge 1991.

Subimago. CUTICULAR COLORATION: Cuticle with light brown and colorless areas. Head colorless, only antennae slightly tinged with pale brownish. Pronotum with light brown and colorless areas. Mesonotum at most light brownish, medioparapsidal suture colorless, other sutures darkened ( Fig. 143 ). Thoracic pleura with light brown and colorless areas; lateropostnotal crest narrowly colored by dark brown, diffusively bordered by light brown ( Fig. 144 ). Base of wing light brown, other wing cuticle lighter. Legs at most colorless; distal margin of each tarsal segment narrowly bordered by brown. Abdominal terga and sterna slightly colored with light brownish, microtrichiae brown. Cerci colorless.

HYPODERMAL COLORATION: As in imago ( Kluge 1991).

TEXTURE: On all legs of both sexes all tarsal segments entirely covered by pointed microlepides ( Table 1).

Tarsal spines of subimago and imago. Tarsus of fore leg in female with one apical spine on 3dr tarsomere (in male without apical spines). Tarsus of middle and hind leg of both sexes with 2 apical spines: one sine on 1st+2nd and another spine on initial 3rd tarsomere ( Table 1). Other characters of imago as described by Kluge 1991.

Dimension. Fore wing length 4–6 mm.

Distribution. Cuba.

Figures

FIGURES 136 – 138. Cloeodes spp. 136, Cloeodes auwe, subimaginal sternum IX with styliger and gonostyli developing under larval cuticle (gonovectes shown by interrupted line; at left half gonovectal and gonostylar muscles shown by interrupted lines, hypodermal pigment shown by dots; median styligeral muscle shown by interrupted lines). 137 – 138, Cloeodes superior: 137, exuviae of subimaginal genitals; 138, genitals of male imago (gonovectes shown by interrupted line; at right half gonovectal and gonostylar muscles shown by interrupted lines; median styligeral muscle shown by interrupted lines, areas of anterior attachment of median paraproctal muscles shown by dotted lines). Abbreviations: 1, 2, 3, segments of gonostylus; m. IX-X, areas of anterior attachment of median paraproctal muscles (i. e., muscles going from sternum IX to common base of paraprocts); m. sg, median sterno-styligeral muscle; usg, unistyliger.

FIGURES 139 – 144. Cuban species of Cloeodes. 139 – 141 and 143 – 144, Cloeodes superior; 142, Cloeodes inferior. 139 – 140, left and right mandibles; 141, left mandible of another specimen; 142, apex of maxilla; 143 – 144, subimaginal exuviae: 143, half of mesonotum; 144, postsubalar sclerite and lateropostnotal crest. Abbreviations: ds 1, ds 2, ds 3, dentisetae.

FIGURES 6 – 12. Cloeodes vibratorius sp. n., larva. 6, head of female larva; 7, maxilla; 8, apex of maxilla; 9 – 11, left and right mandibles; 12, labrum (apical setae not shown) (10, 12, holotype). Abbreviations: ds 1, ds 2, ds 3, dentisetae.