Campylopus savannarum (C.M.) Mitt.

P. A. Florschütz, 1964, Flora of Suriname, volume VI, part I, Leiden: Brill : 78-81

6. Campylopus savannarum (C.M.) Mitt. Journ. Linn. Soc. London Bot 12:35. 1869.

Dicranum savannarum C.M. Syn. Musc. 2: 596. 1851.

Type specimen: Kegel s.n., Suriname (BM, L).

The type specimens from the Kegel collection are preserved at Goettingen. However, the type specimen of C. savannarum is not present. Instead there is a specimen Hostmann 605A, which is not cited by Miiller in his original publication of the species. In the British Museum and in Leiden there are specimens with the label: " Dicranum savannarum mihi / Surinam, in savannis arenosis prope Mariearenosis paston in Saramaccam superiorem/ Kegel Gt. Hb. Müller". Müller did not describe the species in his paper on the Kegel collection, but in the Supplement of his Synopsis. Apparently he did not return the material to Goettingen and it has probably been destroyed in Berlin with the Müller herbarium. The above mentioned collections seem to be part of the original material.

Yellowish to dark green plants, growing in loose or compact mats. Stems densely reddish-tomentose, irregularly branched, up to 6 cm long, equally foliate. Leaves erect and appressed or spreading and rigid or flexuous when dry, erect-spreading when moist, sometimes more or less homomallous, especially at end of stem, linear-lanceolate or lanceolate, concave, apex acute or acuminate, 4-7 mm long and 0,5-0,9 mm wide, margin sometimes inflexed above, serrate up to 1/4 down. Costa strong, short- to long-excurrent, the excurrent part serrate all around and sometimes hyaline, forming an arista, on back with prominent ridges and serrate in upper part, in cross-section showing a dorsal and a ventral row of stereids. Cells along the costa, just above the alar group, rectangular, 15-55 µ long and 15-20 µ wide, walls not or very slightly and inconspicuously pitted, smaller and quadrate towards margin and often transversely elongate at extreme margin; towards apex gradually smaller and more irregular, from linear and elongate-rhomboidal to transversely rhomboidal or oval, 12-30 µ long and 3-9 µ wide. Alar cells variable, often reddish-brown, inflated and conspicuous, sometimes inconspicuous. Sporophyte unknown. (Fig. 23)

Distribution Central America, Guianas.

Two subspecies may be distinguished:

1 a. Leaves spreading when dry, widely spreading with recurved tips when moist, flexuous, linear-lanceolate, apex acute, usually 6-7 mm long and 0,5-0,65 mm wide ssp. savannarum

b. Leaves usually erect when dry, rigidly appressed to the stem, spreading but never recurved or flexuous when moist, lanceolate, apex often more or less acuminate, usually 4-5 mm long and 0,5-0,9 mm wide. ssp. bartlettii

ssp. savannarum.

The alar cells in this subspecies are very conspicuous, brown. The habit, with flexuous, spreading leaves,is characteristic.

On sand in savannas in light shady places. Not common. Near J densavanna and near Mariepaston on Saramacca River. Also in British Guiana. Not collected in French Guiana.

Spsecimens examined: Kegel s.n. (BM, L, type specimen); Lindeman 4032. Without locality: Hostmann 605A (BM, GOET, L); s.n. (PC-MO); without collector or locality (BM). British Guiana: Appun 583 P-P (NY, K); Bartlett s.n. (NY, 3.5 C. sprucei ) Pearce s.n. (NY, K, as C. sprucei ).

ssp. bartlettii (Bartr.) Florsch. comb. nov.

Campylopus bartlettii Bartr. Journ. Wash. Ac. Sci. 22: 477. 1932.

Type specimen: Bartlett 12973, British Honduras (NY)

This subspecies is characterized by the strictly appressed leaves when dry, never flexuous or recurved when moist. The alar cells are often inconspicuous and the excurrent part of the costa is sometimes hyaline. On rocks. Apparently confined to higher altitudes in the interior part of the country Gran Rio (400 m), Kassikassima Mountain (400 m), Voltzberg (230 m). Also in British Guiana. Not collected in French Guiana.

Specimens examined: Tresling 48513; Pulle 240; Rombouts s.n.; s.n.; Mennega 55; Schulz 7762, 7772. British Guiana: Appun 583 (NY, 2 specimens, Genskes one as C. detonsum and as C. sprucei ); A. C. Smith 3645 (NY, K, U), 3656 (NY).

µ µ

C. sprucei Mitt. Journ. Linn. Soc. London Bot. 12: 81. 1869; type specimen: Spruce 59, Brazil (NY), seems to differ only in the distinctly pitted, basal cells.