Tomopterna tuberculosa (Boulenger, 1882)

Conradie, Werner, Keates, Chad, Verburgt, Luke, Baptista, Ninda L. & Harvey, James, 2023, Contributions to the herpetofauna of the Angolan Okavango- Cuando-Zambezi river drainages. Part 3: Amphibians, Amphibian & Reptile Conservation (e 325) 17 (1), pp. 19-56 : 49

Tomopterna tuberculosa (Boulenger, 1882)

Rough Sand Frog ( Fig. 37; Map 36)

Material (5 specimens): PEM A12632–3, roadside ditch 10 km SW of Cuito, -12.44815° 16.88118°, 1,743 m asl; PEM A12681, INBAC: WC-5235, Camp 3, Malova Village, Mipanha River, -14.09140° 16.41476°, 1,553 m asl; PEM A13792–3, Dam/Hydroplant on Rio Cuquema, -12.42556° 16.81856°, 1,640 m asl. Description: Medium sized frog; short snout; tympanum clearly visible; elevated ridge from behind eye to arm; reduced webbing; inner metatarsal tubercle large; subarticular tubercles single. Dorsum brown with near-symmetrical dark brown markings with thin white border; dark brown interorbital bar; no light vertebral line. Adult females (n = 2) measured 45.4–46.3 (45.8) mm (largest female: PEM A12681); adult males (n = 4) varied from 31.9– 36.5 (32.3) mm (largest male: PEM A12633). Habitat and natural history notes: This species was only found within the western side of the study area, associated with open and rocky habitats. Comments: This species occurs mostly in central and western Angola, with isolated records in the east ( Marques et al. 2018). Elsewhere, it is known from northern Namibia and western Zambia to northern Tanzania and south to Zimbabwe ( Channing and Rödel 2019).