Dardanus janethaigae, Parente & Hendrickx, 2009

Parente, Manuel Ayón & Hendrickx, Michel E., 2009, 2323, Zootaxa 2323, pp. 1-71 : 40-52

Dardanus janethaigae n. sp.

( Figs. 6D, 13D, 20–25, 37A–E)

Dardanus sinistripes .— Ayón Parente, 1997: 45 (in part, see material examined and additional material). Hendrickx, 1990; 41 (in part, see additional material); 1994: 27–31 (in part, see material examined and additional material). Hendrickx et al., 1997a: 9 (in part, see additional material).

Material examined. Type material. Holotype male (SL 15.30 m), off Rocas Hermanas , Sinaloa, Mexico, 23º11’07”N, 106º20’21”W, 13 Jun 1996, 6 m, gill net, EMU-4960 . Paratypes: 1 ovigerous female (SL 11.90 mm), off Hermanas Rocks, Sinaloa, Mexico, 23º11’07”N, 106º20’21”W, 13 Jun 1996, 6 m, gill net, EMU- 8471 ; 4 males (SL 4.50–7.80 mm), 1 female (SL 4.30 mm), 3 juvs., off Punta Piaxtla , Sinaloa, Mexico, 23º34’N, 106º57’W, SIPCO I, R / V “El Puma”, 24 Apr 1981, 66 m, trawl, EMU-203 ; 3 males (SL 9.60–12.00 mm), 1 ovigerous female (SL 12.90 mm), off El Delfin “Oceanica”, Sinaloa, Mexico, 23º20’15”N, 106º29’26”W, 11 Aug 1995, gill net, EMU-4775 ; 1 male (SL 4.22 mm), ATLAS II 1 A–01, 21º15´36”N, 105º16´30”W, 20 m, EM-10629; 2 males (SL 2.30–4.00 mm), 2 females (SL 3.00– 4.20 mm), Gulf of California , 28º28’N, 112º04’30”W, 23 Mar 1889, “Albatros” R / V , trawl, 53 m, USNM-265359; 2 males (SL 8.60–8.70 mm), 1 ovigerous female (SL 8.20 mm), Guaymas Area , Sonora, Gulf of California, 19 Aug 1982, 3 m, LACM CR 1982 - 003.1 ; 1 male (SL 10.63 mm), San Agustin Beach ( 30 mi N of San Carlos ), Sonora, Mexico, 21 Nov 1982, UAZ-1416 .

Additional material. 12 juvs., off Teacapán, Sinaloa, Mexico, 22º17’N, 106º10’W, SIPCO I R / V “El Puma”, 23 Apr 1981, 61 m, trawl, EMU-200A; 2 males (SL 6.70–7.56 mm), 2 NS, off Punta de Mita Nayarit, Mexico, 20º53’N, 105º27’W, CORTES 1 R / V “El Puma”, 23 Mar 1985, 48 m, trawl, EMU-207C; 2 NS, north of Tiburón Island, Sonora, Mexico, 29º11’N, 112º31’W, CORTES 3 R / V “El Puma”, 02 Aug 1985, 77 m, trawl, EMU-209 ; 1 male (SL 4.50 mm), off San Miguel Cape , BC, Mexico, 28º10’N, 112º47’W CORTES 2 R / V “El Puma”, 13 Mar 1985, 27 m, trawl, EMU-218A ; 2 males (SL 6.80–8.50 mm), 2 females (SL 4.00– 5.90 mm), and 4 NS, off San Miguel Cape , BC, Mexico, 28º6’N, 112º47’W, CORTES 3 R / V “El Puma”, 01 Aug 1985, 25 m, trawl, EMU-218B ; 1 female (SL 5.50 mm), off Santa María Bay , Sinaloa, Mexico, 25º2’N, 108º31’W, CORTES 1 R / V “El Puma”, 03 May 1982, 28 m, trawl, EMU-216A ; 1 male (SL 6.40 mm), 1 female (SL 5.50 mm), off Santa María Bay , Sinaloa, Mexico, 24º56’N, 108º44’W, CORTES 2 R / V “El Puma”, 10 Mar 1985, 64 m, trawl, EMU-216B ; 1 male (SL 3.60 mm), off Gorda Bank , BCS, Mexico, 23º5’ N, 109º31’ W, CORTES 1 R / V “El Puma”, 13 May 1982, 56 m, trawl, EMU-214A ; 1 male (SL 5.30 mm), off Santa Inés Bay , BCS., Mexico, 26º58´N, 111º53´W, CORTES 2 R / V “El Puma”, 19 Mar 1985, 68 m, trawl, EMU-214B ; 2 females (SL 3.70–9.10 mm), off Santa Inés Bay , BCS, Mexico, 26º58’N, 111º53’W, CORTES 2 R / V “El Puma”, 19 Mar 1985, 68 m, trawl, EMU-214C ; 1 male (SL 3.80 mm), off Fuerte River , Sinaloa, Mexico, 25º46’N, 109º35’W, CORTES 2 R / V “El Puma”, 20 Mar 1985, 97 m, trawl, EMU-214D ; 3 females (SL 4.80–7.30 mm), off Punta Arboleda , Sonora, Mexico, 26º52’N, 110º3’W, CORTES 2 R / V “El Puma”, 12 Mar 1985, 24 m, trawl, EMU-217A ; 1 male (SL 6.90 mm), off Punta Arboleda , Sonora, Mexico, 26º52’N, 110º1’W, CORTES 3 R / V “El Puma”, 31 Jul 1985, 22 m, trawl, EMU-217B ; 1 female (SL 4.60 mm), off Fuerte River , Sinaloa, Mexico, 25º40’N, 109º28’W, CORTES 2 R / V “El Puma”, 14 Mar 1985, 31 m, trawl, EMU-213A ; 1 female (SL 4.50 mm), off Fuerte River , Sinaloa, Mexico, 25º43’N, 109º29’W, CORTES 3 R / V “El Puma”, 08 Aug 1985, 23 m, trawl, EMU-213B; 11 NS, off Punta Arboleda, Sonora, Mexico, 26º51’N, 110º5’W, CORTES 3 , R / V “El Puma”, 05 May 1982, 53 m, trawl, EMU-211 ; 2 males (SL 2.50–3.50 mm), 2 females (SL 3.30–3.90 mm), 1 NS, South of Carmen Island , BCS, Mexico, 25º34’N, 111º8’W, CORTES 1 R / V “El Puma”, 04 May 1982, 55 m, oyster dredge, EMU-212A ; 3 females (SL 5.10–6.70 mm), South of Carmen Island , BCS, Mexico, 25º58’N, 111º07’W, CORTES 1 R / V “El Puma”, 04 May 1982, 50 m, oyster dredge, EMU-212B ; 1 male (SL 6.97 mm), North of Tiburón Island , Sonora, Mexico, 29º12’N, 112º31’W, CORTES 1 R / V “El Puma”, 07 May 1982, 75 m, trawl, EMU-212C ; 1 female (SL 6.10 mm), North of Tiburón Island , Sonora, Mexico, 29º20’N, 112º26’W, CORTES 1 R / V “El Puma”, 07 May 1982, 47 m, trawl, EMU-212D;; 3 males (SL 7.60–13.00 mm), 1 female (SL 11.20 mm), off San Miguel Cape , BC, Mexico, 28º09’N, 112º46’W, CORTES 1 R / V “El Puma”, 06 May 1982, 34 m, oyster dredge, EMU-231 ; 1 male (SL 9.80 mm), 2 females (SL 9.35–11.20 mm), 4 ovigerous females (SL 6.50–6.90 mm), off River , Sonora, Mexico, 27º38’N, 110º44’W, GUAYTEC II R / V “El Puma”, 01 Aug 1987, 82 m, trawl, EMU-229 ; 5 males (SL 9.20–12.80 mm), 1 female (SL 11.40 mm), 2 ovigerous females (SL 10.50–12.10 mm), Los Puentes Cuates, Sinaloa, Mexico, 28 Sep 1996, gill net, EMU-4934B ; 1 male (SL 6.40 mm), off Mazatlán , Sinaloa, Mexico, 23º8’N, 106º25’W, SIPCO I R / V “El Puma”, 25 Apr 1981, 27 m, trawl, EMU-3835A; 1 NS, off Estero Tastiota, Sonora, Mexico, 28º16’N, 111º36’W, CORTES 3 R / V “El Puma”, 06 Aug 1985, 57 m, trawl, EMU-3839D ; 2 males (SL 4.50 mm), off Punta San Marcial , BC, Mexico, 25º33’N, 110º58’W, CORTES 3 R / V “El Puma”, 30 Jul 1985, 41 m, oyster dredge, EMU-3839A ; 1 female (SL 5.40 mm), off Punta Arboleda , Sonora, Mexico, 26º56’N, 110º5’W, CORTES 1 R / V “El Puma”, 05 May 1982, 28 m, trawl, EMU-3841A ; 2 males (SL 4.20–7.00 mm), 1 female (SL 4.10 mm), off Cabo San Miguel, BC, Mexico, 28º9’N, 112º46’W, CORTES 1 R / V “El Puma”, 06 May 1982, 34 m, oyster dredge, EMU-3841B; 6 juvs., off Tepoca Bay , Sonora, Mexico, 30º3’N, 112º55’W, 100 m, trawl, EMU-3841C; 3 NS, off Estero Tastiota, Sonora, Mexico, 28º16’N, 111º31’W, CORTES 1 R / V “El Puma”, 11 May 1982, 49 m, trawl, EMU-3841D; 1 NS, off Río Baluarte, Sinaloa, Mexico, 22º39’N, 106º14’W, BIOCAPESS VI R / V “El Puma”, 28 Jun 1992, 79 m, trawl, EMU-4830B ; 2 females (SL 4.10–4.60 mm), Novillero , Nayarit, Mexico, 22º22’N, 105º56’W, BIOCAPESS VI R / V “El Puma”, 29 Jun 1992, 44 m, trawl, EMU-4830C ; 1 male (SL 6.26 mm), off Puerto Madero , Chiapas, Mexico, 14º37’N, 92º57’W, CEEMEX P7 R / V “El Puma”, 12 May 1992, 118 m, trawl, EMU- 4831D ; 1 male (SL 9.80 mm), Venados Island , Mazatlán, Sinaloa, Mexico, 17 May 1996, EMU-4959 ; 1 female (NM), Rocas Hermanas , 23º11’06”N, 106º26’22”W, 22 Mar 2005, Van Veen dredge, 4 m, EMU-8472 ; 1 male (SL 11.60 mm), Rocas Hermanas , Mazatlán, Sinaloa, Mexico, 23º11’02”N, 106º26’24”W, 10 Nov 2007, 6 m, gill net, EMU-8473 ; 2 males (SL 5.30–7.40 mm), off Huatabampo , Sonora, Mexico, 26º47’N, 110º6’W, GUAYEC II R / V “El Puma”, 01 Aug 1987, 85 m, trawl, EMU-223 .

(All specimens in holdings of EM) 1 male (SL 8.24 mm), ATLAS II, 17º07’N, 100º37’W, 20 Apr 1982, 30 m, trawl, EM-3235 ; 1 male (SL 3.58 mm), ATLAS I, stn. 8B-21, 16º13’24” to 16º14’06”N, 98º44’36” to 98º45’36”W, 16 Feb 1982, 40 m, trawl, EM-3251 ; 1 male (SL 11.00 mm), ATLAS III, stn. 5B-0B, 16º36’36” to 16º38’56”N, 99º32’36” to 99º33’30”W, 17 Jan 1983, 57 m, trawl, EM-3272 ; 1 male (SL 6.50 mm), 3 NS, ATLAS III 6 B-04, 16º34’48” to 16º34’54”N, 99º03’36” to 99º04’48”W, 17 Jan 2003, 32 m, EM-3339 .

(All specimens in holdings at Smithsonian, USNM) 1 NS, Gulf of California, 24º11’30”N, 109º55’W, 30 Apr 1888, R / V "Albatross", oyster dredge, 18 m, USNM-265364; 1 female (SL 6.00 mm), Gulf of California , 30º58’30”N, 113º17’15”W, 24 Mar 1889, R / V "Albatross", beam trawl, 20 m, USNM-265361 ; 1 female (3.60 mm), Gulf of California , 25º02’15”N, 110º43’30”W, 17 Mar 1889, R / V "Albatross", oyster dredge, 31 m, USNM-265365 ; 1 female (SL 9.80 mm), San Jose Island , Gulf of California, 24º54’45”N, 110º39’30”W, 1889, R / V "Albatross", stn. 3000, USNM-1076127 ; 1 male, (SL 7.30 mm), San Jose Island , off NW shore, Amortajada Anchorage, Gulf of California, stn. UH 04167, USNM-1076131 ; 2 males, (SL 8.40–9.90 mm), 2 NS, Isabela Island , Nayarit, Mexico, Allan Hancock Expedition, R / V "Velero III ", stn. 277-34, 5 Mar 1934, USNM-1076119 ; 1 female (SL 3.80 mm), Carmen Island , Gulf of California, 15 Dec 1931, 73 m, USNM- 1107136 .

(All specimens in holdings of LACM CR) 1 male (SL 4.0 mm), 4 females (3.30–7.90 mm), stn. 673-37, 14 Mar 1937, 37 m, LACM CR- 1937- 075.2 ; 1 female (SL 11.50 mm), Searcher Expedition 499-1, shore, LACM CR- 1972- 071.1 ; 1 male (SL 7.90 mm), south side Tiburon Island, Gulf of California, stn. P-194-60, 27 Mar 1960, Parker dredge, 12 m, LACM CR- 1960- 005.2 ; 2 females (SL 3.2–4.0 mm), off Robelar, Sinaloa, Gulf of California, stn. VS-BII-30, 12 Mar 1959, 88 m, LACM CR- 1959- 001.4 ; 1 female (SL 9.70 mm), Puerto Refugio, Angel de la Guarda Island, Gulf of California, stn. 1051-40, 27 Jan 1940, 38 m, LACM CR- 1940- 023.2 ; 1 female (5.00 mm), south of San Francisco Island , Gulf of California, stn. 514-36, 24 Feb 1936, 27 m, LACM CR- 1936- 028.1 ; 1 female (SL 7.40 mm), east of San Francisco Island , Gulf of California, stn. 514-36, 24 Feb 1936, shore, LACM CR- 1936- 028.2 ; 3 males (SL 8.60-9.40 mm), 1 female (SL 8.70 mm), off Concepción Bay, Gulf of California, stn. 681-37, 15 Mar 1937, 5 m, LACM CR- 1937- 083.1 ; 2 males (SL 4.00– 7.60 mm), 1 female (4.30 mm), south of Tiburón Island, Gulf of California, stn. 1044-40, 25 Jan 1940, 29 m, LACM CR- 1940- 016.2 ; 1 female (SL 4.00 mm), 2 juvs., San Ignacio Bay , Sinaloa, Gulf of California, stn. 742-37, 31 Mar 1937, 55– 91 m, LACM CR- 1937- 145.1 ; 1 female (SL 6.50 mm), off Concepción Bay , Gulf of California, stn. 680-37, 37 m, LACM CR- 1937- 082.1 ; 1 female (SL 4.50 mm), San Francisco Island , Gulf of California, stn. 653-37, 9 Mar 1937, LACM CR- 1937- 055.1 ; 2 male s (SL 3.50–7.10 mm), 1 juv., off Pulpito Point, Gulf of California, stn. 674-37, 14 Mar 1937, 26 m, LACM CR- 1937- 076.1 ; 3 males (SL 6.30– 9.60 mm), off Punta Piaxtla, Sinaloa, Gulf of California, stn. VS-BII-33, 23º33.3’N, 106º52.6’W, 13 May 1959, 46 m, LACM CR- 1959- 001.5 ; 2 males (SL 5.10–5.50 mm), 1 female (5.80 mm), 2 juvs. (SL 3.20–3.30 mm), off Punta Bahía Kino , Sonora, stn. P-196-60, 27 Mar 1960, 24 m, LACM CR- 1960- 005.3 .

(All specimens in holdings at University of Arizona, UAZ) 1 ovigerous female (SL 4.21 mm), stn. 16-25, 24º16’N, 110º24’W, R / V “Te Vega”, 44–66 m, UAZ-794; 1 male (SL 4.04 mm), NW of San Carlos , Algodones Bay, Guaymas, Sonora, Mexico, 24 Mar 1970, trawl, 55 m, UAZ-806 .

Diagnosis. Merus of third maxilliped with at least 2 spines on ventral margin. Left cheliped short and broad. Setae on outer face of palm of left cheliped long, plumose, usually reaching subdistal margin of the next scale. Scales on the palm of left cheliped large and subrectangular, to subcircular concealed by long plumose setae. Outer lower angle of carpus of left cheliped bearing a strong spine. Upper face of dactyl of left cheliped with 2 rows of strong spine-like tubercles terminating in single or bifid corneous-tipped spine. Outer face of palm of the left cheliped and the lateral face of the left third pereopod dactyl and propodus concealed with long plumose setae. Lateral face of dactyl of left third pereopod with scales on both sides of longitudinal groove, fringed with small corneous-tipped spines and long plumose setae. Corneous-tipped spines along entire ventrolateral margin of the dactyl of the left third pereopod.

Description. Shield ( Fig. 20A, B) 1.00–1.10 times longer than broad; anterior margin between lateral projections and rostrum almost straight to moderately concave; lateral margins convex, somewhat irregular, with small spines or teeth on anterior 1/4 and tufts of long setae; anterolateral margins usually spiny or toothed; dorsal face of shield flat, with tufts of long setae; weakly calcified Y-shaped line present posteriorly; posterior margin rounded. Rostral lobe indistinct or only weakly produced. Lateral projections moderately pronounced, usually with one (ocasionally two) small spine on distal margin and denticles on lateral margin. Posterior carapace lateral elements well calcified, unarmed. Branchiostegites unarmed.

Ocular peduncles ( Fig. 20A) 0.35–0.50 length of shield, stout, slightly compressed in middle, usually bearing 11 bristles on the superior notch; cornea dilated, 1.16–1.25 width of base of ocular peduncle; corneal length 0.37–0.40 of ocular peduncle length. Ocular acicles ( Fig. 20A, B) broad; distal margins each with 5–7 spines, tips often black; separated by 0.33–0.40 width of an acicle. Interocular plate ( Fig. 20A, B) transversally subrectagular with pair of protrusions.

Antenular peduncles ( Fig. 20A,) slender, 1.33–1.50 length of ocular peduncles; ultimate segment with 3 long stiff setae on dorsal margin, penultimate segment with tufts of long stiff setae dorsally; basal segment ( Fig. 20C) with ventromesial distal angle bearing small spines and armed ventrally with row of 4–7 small spines.

Antennal peduncles ( Fig. 20A, D) when totally extended 1.00–1.33 length of ocular peduncles; fifth segment unarmed, with tufts of short setae on dorsal and ventral faces; fourth segment with 1 or 2 small spines or spinules on laterodistal margin and tufts of long setae on ventrolateral margin; third segment with small spine on ventral subdistal margin and tufts of long stiff setae; second segment with dorsomesial distal angle bearing spines, dorsolateral distal angle pronounced, ending in bifid spines, lateral margin unarmed; first segment with 1 small spine on ventrolateral subdistal margin, ventral surface strongly pronounced, rounded. Antennal acicle ( Fig. 20A, B, D) short, not reaching base of corneas, terminating in bifid spine; dorsomesial margin with 5–7 small spines; dorsolateral margin with 1 or 2 small spine on distal third; ventrolateral margin occasionally with 1 or 2 small spines.

Maxillule ( Fig. 21A) with internal lobe bearing 1 proximal seta and 3 or 4 stiff setae or bristles distally. Maxilla ( Fig. 21B) with endopod moderately long, exceeding scaphognathite in distal extension. First maxilliped ( Fig. 22C) with endopod shorter than basal segment. Second maxilliped ( Fig. 21D) without distinguishing characters. Third maxilliped ( Fig. 21E) with basis-ischium incompletely fused; coxa with 1 moderately strong spine on lateral proximal margin accompanied with tufts of long setae; ischium with well developed crista dentata, with 9–11 denticles; ventrolateral margin with strong distal spine; merus with 2 or 3 spines on ventral margin, dorsodistal margin with small spine; propodi and dactyli usually unarmed propodus and dactyl of left maxilliped of male holotype bearing 2 and 1 moderately strong spines on dorsal face, respectively.

Chelipeds vastly unequal, left larger. Left cheliped ( Figs. 6D, 22A) very stout, 1.30–1.40 times longer than wide, proportion and armature generally similar in males and females. Dactyl terminating in large corneous claw; cutting edge with 5 strong molar teeth; outer face ( Fig. 22C) with row of strong spine-like tubercles terminating in a corneous spine or rounded; upper face ( Fig. 22C) with 2 rows of strong spine-like tubercles terminating in single or bifid corneous-tipped spine, occasionally the middle row consists of scalelike tubercles usually terminating in small, rounded or corneous-tipped spines, tubercles fringed anteriorly with tufts of long plumose setae; inner face ( Fig. 22D) with two rows of tufts of stiff setae, the row near upper margin bearing corneous spine-like tubercles. Fixed finger terminating in strong corneous claw; cutting edge with 5 or 6 strong molar teeth. Palm ( Figs. 6D, 22A, B) with outer face strongly convex, with 2 longitudinal rows of 7–9 prominent spines on the upper face, generally a row of smaller spines present between these two rows; upper outer face with several longitudinal, irregular rows of both, spine-like and scale-like tubercles terminating in rounded or corneous spines; outer faces of palm and fixed finger covered with scales; each scale fringed anteriorly with 1–7 small granules or corneous-tipped spines and tufts of long plumose setae which usually reaching the subdistal margin of the next scale; inner lower portion with numerous flattened tubercles bearing 1 or 2 rounded spines; lower margin ( Fig. 22D) with very strong, triangle-shaped corneous spines. Carpus ( Fig. 22A) with upper margin bearing one row of prominent corneous-tipped spines; upper face with several corneous-tipped spines, spines larger near upper margin; outer face with numerous corneous-tipped spines; lower outer faces with corneous-tipped spines on proximal and distal margins; inner face with scarce tufts of short setae, inner distal angle usually with 4 strong or moderately strong spines or teeth and scarce tufts of long setae. Merus with distal margin of the lateral face bearing several corneoustipped spines, larger spine dorsally; dorsal face with tufts of long stiff setae, subdistal margin with short transversal row of small corneous-tipped spines; lateral face with numerous flattened tubercles bearing small corneous spines and tufts of scarce, long, stiff setae; ventrolateral angle with 1 or 2 strong corneous-tipped spines; ventromesial margin crested with 6 teeth, 3 proximal larger than distal. Ischium with moderately strong rounded spines on ventromesial margin.

Rigth cheliped ( Fig. 22E, F) moderately slender, generally setose. Dactyl terminating in large corneous claw; cutting edge with 4 or 5 molar teeth; upper face with 3 irregular longitudinal rows of strong or moderately strong corneous-tipped spines accompanied with tufts of long stiff setae; outer face with longitudinal row of 3–5 corneous-tipped spines accompanied with tufts of long stiff setae. Fixed finger terminating in large corneous claw; cutting edge with 5 or 6 molar teeth. Palm with 2 irregular rows of prominent corneous-tipped spines accompanied by tufts of long stiff setae on upper margin; upper outer face near upper margin with several moderately strong corneous-tipped spines; outer face of palm and fixed finger with numerous flattened tubercles bearing triangle-shaped corneous spines; lower face with 2 irregular longitudinal rows of strong, flattened, triangle-shaped corneous spines accompanied with tufts of long stiff setae ( Fig. 22F). Carpus with 8 or 9 peripheral, strong corneous-tipped spines on upper face; outer and lower outer faces with several flattened tubercles bearing small corneous-tipped spines; ventrolateral angle with 1 strong, corneous-tipped spine; inner lower angle with 3 teeth or rounded spines. Merus with distal margin of lateral face bearing several corneous-tipped spines, larger spine dorsally; dorsal face with tufts of long stiff setae; lateral faces with tufts of stiff setae; ventrolateral angle with 1 strong corneous-tipped spine; ventromesial margin crested with 6 rounded teeth, proximal larger. Ischium with 5 rounded teeth or spines on ventromesial margin.

Second pereopods ( Fig. 23A–D) and right third pereopod ( Fig. 24A–D) generally similar, but armament of dactyls, propodi and carpi somewhat different between second pair and right third; second pair generally more slender than right third; of second pair, left slightly shorter than right. Dactyls 1.30–1.40 (second) or 1.30–1.50 (right third) length of propodi, each terminating in large corneous claw; dorsal surfaces ( Fig. 23C) each with row of flattened tubercles, bearing 2 corneous spines and tufts of long, stiff setae; lateral and mesial faces each with median weak longitudinal groove and two rows of tufts of stiff setae, one in midline and another dorsally, the latter generally accompanied with corneous spines; ventral margins with rows of tufts of long stiff setae, ventrolateral row of left second pereiopod ( Fig. 23A) also with small corneous spines on proximal third, of right third ( Fig. 24A) with corneous spines in each tuft, each row with 5–7 (second) or 2 or 3 (right third) distal, corneous spines. Propodi 1.30–1.40 length of carpi; dorsal faces flattened and very broad (second, Fig. 23D) or comparatively narrower (right third, Fig. 24D), each with 3 irregular longitudinal rows of flattened tubercles bearing 1 or 2 corneous spines and tufts of long stiff setae; lateral faces each with 2 rows of tufts of stiff setae, one in midline and another ventrally, in the left second ( Fig. 23A) both rows of tufts bearing small corneous spines; mesial faces ( Figs. 23B, 24C) each with 2 rows of tufts of stiff setae, one row in midline, another ventrally (right third), both rows submarginal in second pereiopod, upper row generally bearing corneous spines; ventral margin with tufts of long, stiff setae, that on left second bearing corneous spines. Carpi 0.70–0.80 (second) or 0.90–1.00 (right third) length of meri; dorsal faces each with row of corneous-tipped spines increasing in size distally and tufts of scarce long stiff setae, one row of flattened tubercles bearing small corneous-tipped spines running near dorsal margin (second, Fig. 23D), dorsolateral angles each usually bearing 1 small, corneous-tipped spine; lateral faces convex, with weak longitudinal groove flanked ventrally by flattened tubercles bearing corneous spines and tufts of long stiff setae (second, Fig. 23A) or only tufts of setae (right third, Fig. 24A). Meri laterally compressed, each with tufts of very thick setae on dorsal face; lateral and mesial faces with scarce tufts of short setae; ventrolateral distal margin usually with 1 moderately strong corneous-tipped spine; ventral faces with double irregular row of small spines or granules (second) or scarce small spines or granules distally (third) and tufts of long setae.

Left third pereopod ( Figs. 13D, 25A–D) generally very stout. Dactyl 1.40–1.50 length of propodus, very broad, terminating in a strong corneous claw; dorsal face with longitudinal row of flattened tubercles bearing corneous spines and tufts of stiff setae, larger and denser distally; mesial face ( Fig. 25C) convex, with two longitudinal rows of tufts of stiff setae, one in midline and another dorsally, the latter usually bearing corneous spines; lateral face ( Figs.13D, 25A, B) flattened, with deep longitudinal median groove vanishing distally, lined with row of small flattened tubercles fringed distally with long plumose setae; both sides of groove are transverse scales, fringed on their distal edge with small corneous-tipped spines and long plumose setae, scales terminating in strong corneous spine-like tubercles on dorsal and ventral margins of dactyl, spines partially concealed by tufts of long setae ( Figs. 24E, F, 25B); ventral margin with tufts of longs stiff setae and 1 or 2 small corneous spines distally. Propodus 1.30–1.40 length of carpus, very broad; dorsal face ( Fig. 25D) with 2 irregular longitudinal rows of flattened tubercles bearing 1 or 2 strong corneous spines and tufts of long stiff setae, dorsodistal margin with few corneous spines and long stiff setae; lateral face ( Fig. 25A) convex with two rows of scales on both sides of a bare, median area; scales fringed with small corneous-tipped spines and plumose setae on distal edge, setae longer toward edges of propodus. Scales on lower face terminating in strong, corneous-tipped, spine-like tubercles on ventral margin, scales partially concealed by tufts of long plumose setae and scarce long stiff setae, scales on dorsolateral margin terminating in strong tubercles bearing 2 strong, corneous-tipped spines partially concealed by tufts of long, plumose setae and scarce, long stiff setae; mesial face ( Fig. 25C) with two longitudinal row of short stiff setae, one in midline and another ventrally, distal margin with 3 or 4 corneous-tipped spines. Carpus 0.80–0.90 length of merus; dorsal face bearing row of strong, corneous-tipped spines; dorsodistal angle usually bearing 1 moderately strong, corneous-tipped spine; lateral face convex, with weak longitudinal groove flanked ventrally by flattened tubercles bearing small corneous spines and tufts of long stiff setae, laterosubdistal margin spiny; ventral margin usually with 2 small, corneous-tipped spines accompanied by tufts of long stiff setae. Merus laterally compressed, with armament and setation similar to those of right third.

Sternite XII (third peropods) ( Fig. 20E) with anterior lobe rectangular, bearing an elongate protuberance with tuft of long setae anteriorly.

Fourth peropod ( Fig. 20F) subchelate; dactyl with 6–8 corneous, ventral spines on lateral face; propodal rasp well developed; carpus with sharp dorsodistal spine.

Fifth pereopod chelate; rasps of dactyl and propodus well developed.

Male pleon ( Fig. 23E) with second to fifth left pleopods fringed with long setae, each with well developed exopod; single, small, triangular, fleshy membranous protuberance fringed distally with long setae present between fourth and fifth pleopods. Female pleon ( Fig. 23F) with second to fifth left pleopods fringed with long setae; second to fourth triramous; fifth biramous, endopod very small; single, elongate, fleshy membranous protuberance fringed with long setae present between fourth and fifth pleopods.

Uropods ( Fig. 23E, F) strongly asymmetrical, left larger than right; endopods and exopods with well developed rasps.

Telson ( Fig. 20G) with lateral constrictions; marginal area partially calcified; posterior lobes separed by median cleft, left larger than right, with 4 or 5 (left) or 4–6 (right) corneous spines and long setae on terminal margin; anterior lobes with long setae on lateral margins.

Color. In life ( Fig. 26A). Carapace yellowish-pink with purplish-brown stains on middle dorsal and lateral faces; shield purplish-brown with yellowish-pink stains principally on lateral margins and small purples spots dorsally, yellow setae with red tip; ocular acicles yellowish-pink; antennal and peduncular segments purplish-brown; antennal and peduncular flagella orange; ocular peduncles purplish-brown with two light purple bands, one proximal and another subdistal, bands ventrally darker, a tuft of yellow setae on notch of cornea; cornea golden; left cheliped purplish-brown, tubercles purples; merus pinkish-purple with a dark red band distally; color of smaller cheliped similar, with some white tubercles on outer face of palm and red, yellow-tipped, or yellow, red-tipped hairs; cutting edge of fingers white, with a narrow, longitudinal line of red. Walking legs purplish-brown with purples tubercles; ischium and proximal half of merus yellowish-pink. Left third pereopod with tranverse ridges of purple across flattened surface of propodus and dactyl.

Etymology. This species is named in honor to Janet Haig in recognition of her outstanding contribution to the taxonomy of Paguroidea and the countless occasions in which she shared her experience and knowledge on anomuran with one of us (MEH).

Distribution. Throughout the Gulf of California and south of Guerrero, Mexico; 3– 118 m.

Remarks. Dardanus janethaigae n. sp. and D. pilosus are somewhat similar in their general aspect, but the left third pereopod propodus of D. janethaigae ( Fig. 37D) is stouter ( Fig. 37D, I). Marginal setae of scales on lateral face of dactyl usually reach the distal margin of the next scale in D. pilosus , instead of reaching to half of the next scale as in D. janethaigae , and setae on ventrolateral margin of dactyl and propodus are longer in D. pilosus . Sculpture on the outer face of palm of left cheliped are similar in these two species, fringed with long plumose setae reaching subdistal or distal margin of the next scale, but cheliped of D. janethaigae is stouter ( Fig. 37A, F, Table 1). Lower inner face of left cheliped ( Fig. 37G) in D. pilosus bears numerous corneous spine-like tubercles; in D. janethaigae there are only rounded granules ( Fig. 37B). The dactyl of left cheliped is proportionally larger and slender in D. janethaigae ( Fig. 37C) than in D. pilosus ( Fig. 37H). The outer lower angle of carpus of left cheliped in D. janethaigae bears one strong spine, in lieu of a small spine ( D. nudus , D. stimpsoni and D. pilosus ) or a rounded spine ( D. sinistripes and D. magdalenensis ). Ocular peduncles and antennal acicles are proportionally longer in D. pilosus than in D. janethaigae (Table I), and the armature of telson shows some obvious differences between these species ( Fig. 37E, J).