Manota multisetosa, Jaschhof & Hippa, 2005

Jaschhof, Mathias & Hippa, Heikki, 2005, The genus Manota in Costa Rica (Diptera: Mycetophilidae), Zootaxa 1011 (1), pp. 1-54 : 14

Manota multisetosa sp. n. ( Fig. 5 )

Description. Large species.

Male. Head. Antenna with fourth flagellomere 1.7 times as long as wide. Maxillary palpus with third palpomere bearing 3 curved sensilla; fourth palpomere with a parasegment apically.

Thorax. Anterior basalare and laterotergite each bare. Episternum 3 with ventral portion setose. Wing. Length 1.7–1.8 mm. Membrane with a few setae along posterior margin. Sc setose dorsally and ventrally. CuA2 fading away proximally.

Terminalia ( Fig. 5 ). Sternite 9 small, trapezoid, with posterior and lateral margins distinct; posterior margin situated at mid length of gonocoxites, with a tiny notch medially. Ventro­lateral lobes of gonocoxites rounded; interiorly with up to 5 setae on a separate rim. Disto­lateral lobes of gonocoxites absent. Gonocoxites in position I+II with some 7 moderately large setae plus a dense, irregular comb of numerous shorter setae extending posteriorly; in position III+IV with 2 pointed megasetae. Gonostyli short, ovoid; with setae of moderate length, setae on dorsal side strong, directed dorsally and inwards. Parastylar lobes distinct, at some distance from gonostyli, large, bare. Tegmen almost evenly tapering to tip. Hypoproct broadly rounded, with a few long setae disto­laterally.

Diagnostic characters. Among the species with bare laterotergites, it has terminalia with disto­lateral lobes of the gonocoxites reduced; gonocoxites dorsally with a dense comb of setae; gonostyli stout and with large setae dorso­apically; and parastylar lobes very distinct, bare and situated at some distance from the gonostyli. Very similar to M. tapantiensis . (See next species).

Etymology. From Latin, multus, much, and setosus, setose, referring to the dense comb of setae dorsally on the gonocoxites.

Types. Holotype. Male, Costa Rica, Puntarenas province, Corcovado National Park, 600 m S Cerro Rincón , 745 m, 23 April–24 June 2002, by Malaise trap, J. Azofeifa ( INBio sample #70842) . Paratypes. 4 males, same data as the holotype ; 1 male, same locality , but 1 Aug.–25 Sep. 2002 ( INBio sample #71459) ; 4 males, same locality , but La Tigrilla , 600 m, 20 Dec. 2001 – 7 Feb. 2002, by Malaise trap, J. Azofeifa ( INBio sample #67158) ; 1 male, same locality , but 7 Feb.–23 April 2002 ( INBio sample #70848) .

Other material studied. 3 males, Limón province, Hitoy Cerere Biological Reserve, Sendero Espavel , 11 March–1 April 2003, E. Rojas, B. Gamboa & W. Arana ( INBio samples #73470, 73627 and 73628) ; 4 males, same locality, but 18 Sep.–7 Oct. 2003, E. Rojas, B. Gamboa, W. Arana, M. & C. Jaschhof ; 1 male, Limón province, Pococí, Braulio Carrillo National Park, Estación Quebrada González , 23 April 2002, P. Hanson & C. Godoy ( INBio sample #67599) ; 1 male, Puntarenas province, Cerro Puma 19 June–8 July 2003, M. Moraga, A. Azofeifa & K. Caballero ( INBio sample #74508) ; 1 male, Puntarenas province, Gofito, Jiménez, Corcovado National Park , Cerro Mueller, 20 Dec. 2001 – 7 Feb. 2002, J. Azofeifa ( INBio sample #67160) ; 1 male, same National Park, but Estación Patos, Sendero a Sirena , 5 May–8 June 2001, J. Azofeifa ( INBio sample #64143) ; 1 male, Ala­ juela province, San Carlos, Volcán Arenal National Park, Sendero Pilón , 6–7 Nov. 2000, G. Carballo ( INBio sample #60084) ; 1 male, Alajuela province, Estación Caribe , Sep.– Oct. 2002, W. Flowers ( INBio sample #71759) .

Figures

FIGURE 5. Manota multisetosa sp. n. A: Male terminalia, ventral view (holotype). B: Male terminalia, dorsal view (specimen from Estación Caribe). Scale = 0.1 mm.