PSEUDOCOCCIDAE

Franco, José Carlos, Russo, Agatino & Marotta, Salvatore, 2011, An annotated checklist of scale insects (Hemiptera: Coccoidea) of Portugal, including Madeira and Azores Archipelagos, Zootaxa 3004, pp. 1-32 : 18-21

FAMILY PSEUDOCOCCIDAE

Atrococcus paludinus (Green, 1921)

Distribution: Mainland

References: Kozár & Franco (1995), Ben-Dov et al. (2010), Danzig & Watson (2010).

Balanococcus scirpi (Green, 1921)

Distribution: Mainland

References: Kozár & Franco (1995), Ben-Dov et al. (2010), Danzig & Watson (2010).

Cucullococcus arrabidensis (Neves, 1954)

Distribution: Mainland

References: Neves (1954b, 1956), Ben-Dov (1994), Marotta & Franco (1999), Ben-Dov et al. (2010), Danzig & Watson (2010).

● Delottococcus proteae (Hall, 1937)

Distribution: Madeira

Comments: first record; collected and identified by A.M.F. Aguiar on Protea magnific a x susannae (var. Suzara), in Santo da Serra (Santa Cruz) (19-10-2000); on Protea cynaroides , in Caniço (Santa Cruz) (7-09- 2000) and Santana (22-08-2006); on Leucadendron tinctum , Protea sp., Protea repens , Protea longifolia “Satin Pink”, in Santa (19-10-2000).

Dysmicoccus boninsis (Kuwana, 1909)

Distribution: Madeira

References: Green (1923), Balachowsky (1939), Williams (1970), Vieira et al. (1983), Ben-Dov (1994), CABI (1995b), Ben-Dov et al. (2010), Borges et al. (2008), Danzig & Watson (2010), Pellizzari & Germain (2010).

Dysmicoccus brevipes (Cockerell, 1893)

Distribution: Azores and Madeira

References: Bensaúde (1926a), Balachowsky (1939), Borchsenius (1949), CAB (1972c), Vieira et al. (1983), Ben-Dov (1994), Borges (2005), Ben-Dov et al. (2010), Borges et al. (2008), Danzig & Watson (2010), Pellizzari & Germain (2010).

Dysmicoccus grassii (Leonardi, 1913)

Ditribution: Madeira

References: Williams & Granara de Willink (1992).

Comments: Williams & Granara de Willink (1992) reported (pp. 13) the presence of this species as D. alazon .

Nipaecoccus nipae (Maskell, 1893)

Distribution: Mainland, Azores and Madeira.

References: Bensaúde (1926b), Neves (1935, 1936c), Balachowsky (1939), Seabra (1942), CAB (1966b), Guimarães (1973), Carneiro (1982), Fernandes (1982), Vieira et al. (1983), Ben-Dov (1994), Carvalho & Aguiar (1997), Carvalho et al. (1997), Carvalho (1999), Borges (2005), Ben-Dov et al. (2010), Franco et al. (2006), Borges et al. (2008), Danzig & Watson (2010), Pellizzari & Germain (2010).

● Paracoccus leucadendri Mazzeo & Franco

Distribution: Mainland and Madeira

References: Mazzeo et al. (2009), Ben-Dov et al. (2010).

Comments: although described from specimens collected in Portugal, it is possibly native to Southern Africa ( Mazzeo et al., 2009); new record for Madeira, collected and identified by A.M.F. Aguiar on Leucadendron sp. in Ribeirinha (Camacha) (7-05-1997).

Phenacoccus latipes Green, 1923

Distribution: Madeira

References: Green (1923), Balachowsky (1939), Vieira et al. (1983), Ben-Dov (1994), Ben-Dov et al. (2010), Borges et al. (2008), Danzig & Watson (2010).

Phenacoccus madeirensis Green, 1923

Distribution: Mainland and Madeira

References: Green (1923), Balachowsky (1939), Vieira et al. (1983), Ben-Dov (1994), Carvalho (1999), CABI (2000), Ben-Dov et al. (2010), Borges et al. (2008), Danzig & Watson (2010), Pellizzari & Germain (2010).

Comments: the presence of this mealybug in Mainland reported by Danzig & Watson (2010) is confirmed by us: specimens collected by Elisabete Figueiredo on Capsicum annuum under greenhouse, in Silveira (Torres Vedras) (30-7-2009).

Phenacoccus monieri Balachowsky, 1939

Distribution: Madeira

References: Balachowsky (1939), Vieira et al. (1983), Ben-Dov (1994), Ben-Dov et al. (2010), Borges et al. (2008), Danzig & Watson (2010).

● Phenacoccus peruvianus Granara de Willink, 2007

Distribution: Mainland and Madeira

References: Beltrà et al. (2010)

Comments: first record for Madeira; collected on Bougainvillea sp. in Funchal, by A.M.F. Aguiar (29-05- 2001; 12-06-2002), by the Phytossanitary Services (2-07-2001), and by A. Azevedo (30-08-2001); in Câmara de Lobos, by A.M.F. Aguiar (22-08-2002), and by V. Balmés (10-11-2010); and in Porto Santo, by A.M.F. Aguiar (25-07-2001). All specimens were identified by A.M.F. Aguiar, except those collected in 2010, whose identification was carried out by J.-F. Germain.

● Phenacoccus yerushalmi Ben-Dov, 1985

Distribution: Mainland

Comments: first record; collected by Luis Silva on Pinus sp. in Costa da Caparica (3-06-1995).

Planococcus citri (Risso, 1813)

Distribution: Mainland, Azores and Madeira

References: Almeida (1898b), Lindinger (1912), Green (1923), Bensaúde (1924, 1926c, 1927a, 1927b), Seabra (1906, 1926, 1930a, 1930b, 1930c, 1941), Almeida (1934, 1935), Neves (1936c, 1952, 1954 a), Balachowsky (1939), Borchsenius (1949), Guimarães (1973), Carneiro (1982), Fernandes (1982, 1992), Vieira et al. (1983), Franco & Carvalho (1990), Schanderl et al. (1995), Carvalho et al. (1997), Franco & Marotta (1999), CABI (1999b), Carvalho (1994, 1999), Borges (2005), Ben-Dov et al. (2010), Franco et al. (2000, 2006), Borges et al. (2008), Danzig & Watson (2010), Pellizzari & Germain (2010).

● Planococcus ficus (Signoret, 1875)

Distribution: Mainland, Azores and Madeira

References: Almeida (1898b), Seabra (1906, 1926, 1941), Almeida (1934, 1935), Ezzat & McConnell (1956), Carneiro (1982), Franco et al. (2000), Borges (2005), Ben-Dov et al. (2010), Danzig & Watson (2010). Comments: first record for Madeira; collected by António Miguel Franquinho Aguiar, on Vitis vinifera, in Encruzilhada (S. Vicente) (13-11-1995).

* Planococcus kraunhiae (Kuwana, 1902)

Distribution: Madeira

References: Balachowsky (1939), Vieira et al. (1983).

Comments: Balachowsky (1939) collected this species on Ficus carica and reported that, apparently, it was not differentiable from Pseudococcus citri Risso v. coleorum Marchal. Vieira et al. (1983) did not find it in the Island. This is possibly a misidentification of Planococcus citri (Risso) .

Planococcus minor (Maskell, 1897)

Distribution: Azores and Madeira

References: Borges (2005), Borges et al. (2008).

● Planococcus vovae (Nasonov, 1909)

Distribution: Mainland

Comments: first record; collected by the first author, on Cupressus sp., in Monsanto (Lisbon) (2-10-1995, 8- 11-1995) and Setúbal (29-05-1995) and by Filomena Caetano, on Chamaecyparis sp., in Restelo (Lisbon) (6- 11-1995).

Pseudococcus calceolariae (Maskell, 1879)

Distribution: Mainland, Azores and Madeira

References: Franco & Carvalho (1990), Carvalho (1994), Carvalho et al. (1997), Franco & Marotta (1999), Franco et al. (2000, 2006), Ben-Dov et al. (2010), Borges et al. (2008), Danzig & Watson (2010), Pellizzari & Germain (2010).

Pseudococcus cimensis Green, 1923

Distribution: Madeira

References: Green (1923), Vieira et al. (1983), Ben-Dov (1994), Ben-Dov et al. (2010), Borges et al. (2008), Danzig & Watson (2010).

Pseudococcus comstocki (Kuwana, 1902)

Distribution: Madeira

References: Balachowsky (1939), Borchsenius (1949), Vieira et al. (1983), Ferreira & Guimarães (1984), Ben-Dov (1994), Ben-Dov et al. (2010), Borges et al. (2008), Danzig & Watson (2010), Pellizzari & Germain (2010).

Pseudococcus longispinus (Targioni Tozzetti, 1867)

Distribution: Mainland, Azores and Madeira

References: Almeida (1898b), Seabra (1906, 1930a, 1930c, 1941), Lindinger (1912), Green (1923), Bensaúde (1927b), Neves (1936c, 1952, 1954 a), Balachowsky (1939), Borchsenius (1949), CAB (1958b), Guimarães (1973), Carneiro (1982), Fernandes (1982, 1992), Vieira et al. (1983), Franco & Carvalho (1990), Ben-Dov (1994), Carvalho (1994, 1999), Kozár & Franco (1995), Schanderl et al. (1995), Carvalho & Aguiar (1997), Carvalho et al. (1997), Franco & Marotta (1999), Franco et al. (2000, 2006), Borges (2005), Ben-Dov et al. (2010), Borges et al. (2008), Danzig & Watson (2010), Pellizzari & Germain (2010).

* Pseudococcus maritimus (Ehrhorn, 1900)

Distribution: Madeira

References: Balachowsky (1939), Vieira et al. (1983)

Comments: reported by Balachowsky (1939) from specimens collected at Funchal, Câmara de Lobos and S.Vicente, on several ornamental plants and Trifolium sp., in 1936. According to the present knowledge on the Pseudococcus maritimus complex ( Miller et al. 1984, Gimpel & Miller 1996) this is possibly a misidentification of P. viburni .

Pseudococcus viburni (Signoret, 1875)

Distribution: Mainland, Azores and Madeira

References: Miller et al. (1984), Franco & Carvalho (1990), Fernandes (1993), Carvalho (1994, 1999), Kozár & Franco (1995), Gimpel & Miller (1996), Carvalho et al. (1997), Franco & Marotta (1999), Franco et al. (2000, 2006), Borges (2005), Ben-Dov et al. (2010), Borges et al. (2008), Danzig & Watson (2010), Pellizzari & Germain (2010).

Rhizoecus martini Kozár & Konczné Benedicty, 2007 Distribution: Madeira

References: Kozár & Konczné Benedicty (2007), Ben-Dov et al. (2010).

Ripersiella sasae (Takagi & Kawai, 1971) Distribution: Madeira

References: Kozár & Konczné Benedicty (2007).

Saccharicoccus sacchari (Cockerell, 1895)

Distribution: Madeira

References: Green (1923), CAB (1977b), Vieira et al. (1983), Ben-Dov (1994), Ben-Dov et al. (2010), Borges et al. (2008), Danzig & Watson (2010).

Seabrina cistorum Neves, 1943

Distribution: Mainland

References: Neves (1943), Ben-Dov (1994), Ben-Dov et al. (2010), Danzig & Watson (2010).

Trionymus perrisii (Signoret, 1875)

Distribution: Mainland

References: Kozár & Franco (1995), Ben-Dov et al. (2010), Danzig & Watson (2010).

Vryburgia amaryllidis (Bouché, 1837)

Distribution: Mainland

References: Kozár & Franco (1995), Ben-Dov et al. (2010), Danzig & Watson (2010).