Dadagulella ecclesiola, Rowson & Tattersfield, 2013

Rowson, Ben & Tattersfield, Peter, 2013, Revision of Dadagulella gen. nov., the “ Gulella radius group ” (Gastropoda: Streptaxidae) of the eastern Afrotropics, including six new species and three new subspecies, European Journal of Taxonomy 37, pp. 1-46: 16

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Dadagulella ecclesiola

sp. nov.

Dadagulella ecclesiola   sp. nov. Figs 28 View Figs 27-35 , 49 View Figs 43-50 , 56 View Figs 51-64 , 84 View Fig ; Table 1 View Table 1


From Latin ‘ ecclesiola   ’, feminine, diminutive of ‘church’ (or ‘the church’); used arbitrarily to distinguish the species from D. minareta   sp. nov. with which it occurs.

Type material examined

TANZANIA: holotype NMW. Z.2003.001.00015: 1 ad., Kimboza Forest Reserve (7.01° S, 37.78° E), Uluguru Mts, Morogoro District, lowland forest on limestone, approx. 300 m alt., leg. BR, PT, MBS, & CFN, 5 Feb. 2003. GoogleMaps   Paratypes NMW. Z.2003.001.00016: 8 ads, data as holotype. Paratype MRAC. MT.803794: 1 ad., data as holotype. Paratype NHMUK.20120259: 1 ad., data as holotype. Paratype NMK: 1 ad., data as holotype. Paratype NMT: 1 ad., data as holotype. Paratype NMSA. L 8692/ T 3061: 1 ad., data as holotype. Paratype RMNH. MOL.288089: 1 ad., data as holotype.

Other material examined

TANZANIA: NMW. Z.2003.001.00017: 1 ad., data as holotype, sequenced by Rowson et al. (2010 a) as “ Gulella cf. browni Uluguru   ”. NMW. Z.2003.001.00018: 3 ads in poor condition, 3 juvs, data as holotype.


SHELL ( Figs 28 View Figs 27-35 , 49 View Figs 43-50 , 56 View Figs 51-64 ). Small to medium-sized (3.20 - 3.70 mm high x 1.80 - 1.90 mm wide), of 5.5 - 7.0 whorls. Ovate-acuminate, spire narrowly accuminate (spire angle 53 - 65°). Apex pointed. Embryonic whorls smoothly granulate. Later whorls with relatively coarse, often sinuous ribs (8 - 14 per mm on penultimate whorl). Sutures deep. Umbilicus closed. Peristome complete. Outer palatal surface of aperture with a depression, often furrow-like, corresponding to the palatal tooth. Dentition 4-fold, consisting of: one strongly oblique parietal tooth, V-shaped when shell turned to the left; one large slablike palatal tooth, forming a narrow, horizontal parieto-palatal sinus; and a deep-set columellar baffle. A basal denticle is also present, presumably in all specimens, but is partly or completely hidden by the palatal tooth which occludes much of the aperture. The denticle is visible when the shell is turned to the right ( Fig. 56 View Figs 51-64 ). Juvenile shells not known with certainty: an individual from Kimboza ( Fig. 49 View Figs 43-50 ), with dentition like that of a juvenile D. r. radius   comb. nov., might belong to this species. Anatomy unknown.

Range and habitat

In forest at the type locality in the eastern Tanzanian lowlands.


This species has much simpler dentition than D. minareta   sp. nov. (which also occurs at Kimboza), and is usually smaller, with straight rather than sinuous ribs. Its dentition is more like that of D. r. radius   comb. nov. (which again also occurs at Kimboza; Fig. 16 View Figs 6-26 ) and D. minuscula minuscula ( Morelet, 1877)   comb. nov. but D. ecclesiola   sp. nov. lacks the shallow columellar tooth, has a hidden or partly hidden basal denticle, and has a much narrower parieto-palatal sinus. It further differs from D. minuscula minuscula   comb. nov. in having stronger ribs.


Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet


Harvard University - Arnold Arboretum


Naturhistorisches Museum, Wien


Universität Zürich


Musée Royal de l’Afrique Centrale


Mus. Tinro, Vladyvostok


Natural History Museum, London


National Museums of Kenya


KwaZulu-Natal Museum


Nationaal Herbarium Nederland, Leiden University branch


Tavera, Department of Geology and Geophysics


National Museum of Natural History, Naturalis


Universidad Nacional Agraria La Molina