Eurytenes Foerster, 2011

Walker, Andrea K. & Wharton, Robert A., 2011, A review of New World Eurytenes s. str. (Hymenoptera, Braconidae, Opiinae), Journal of Hymenoptera Research 20, pp. 23-46 : 26-27

Genus Eurytenes Foerster s. str.

Eurytenes Foerster 1862: 259. Type species Opius abnormis Wesmael 1835 by original designation and monotypy.

Description.

Head. Antenna filiform, longer than body. Frons, vertex, and temple smooth, shiny; frons bare, vertex and upper temple nearly so. Labrum exposed. Clypeus weakly to distinctly protruding in profile. Malar sulcus deeply impressed. Mandible gradually to somewhat more abruptly widening from apex to base, carinate ventrally over most of basal half, never with distinct basal tooth as in Opius s. str. Maxillary palp longer than head, reaching mid coxa. Occipital carina present laterally, extending dorsal-medially at least to level of inner eye margin, broadly absent mid-dorsally; widely separated from hypostomal carina at base of mandible.

Mesosoma. Pronotum dorsally with narrow, transverse, crenulate sulcus extending continuously along lateral pronotum to ventral corner; weak to deep median pit interrupting sulcus dorsally. Mesoscutum in profile with anterior declivity very slightly concave, nearly vertical; notaulus present on anterior portion of mesoscutal disc, angled laterally at anterior end, laterally-directed portion carinate along anterior margin; notaulus continuous with weakly to distinctly crenulate impression bordering lateral mesoscutal margin, the impression extending posteriorly at least to level of tegula; midpit of mesoscutum well-developed, discrete. Scutellar sulcus ( Fig. 31 ) narrow, though not exceptionally so, 3-4 times wider than mid-length, crenulate, with numerous closely - spaced ridges. Mesopleuron smooth, shiny, posterior margin not crenulate; precoxal sulcus distinctly impressed, crenulate. Propodeum with large median areola, variously obscured by sculpture.

Wings. Slightly more than twice as long as wide. Stigma long, very narrow, nearly parallel-sided basally, widening distally; thickest part of stigma twice maximum width of proximal half. Radial cross-vein (r) thickened, weakly to strongly bowed anteriorly, arising from extreme base of stigma, nearly in line with 3RSa; RS+M weakly to distinctly sinuate; second submarginal cell nearly parallel-sided, not or only very weakly converging distally; m-cu nearly always postfurcal, entering second submarginal cell; 2CUa distinct, shorter than 2cu-a. Hind wing with both RS and M distinct nearly to wing margin, usually nebulous: very weakly pigmented; m-cu varying from indistinct in smaller individuals to present in larger individuals as a weakly pigmented impression extending more than half way to wing margin.

Legs. Hind femur slender, dorsal surface uneven, somewhat bilobed.

Metasoma. Petiole with distinct dorsope; dorsal and lateral surface densely striate to strigose, largely obscuring dorsal carinae except at base; spiracle of T1 located at or slightly posteriad mid-length; sternite short but distinct, extending 0.4-0.5 distance between base of T1 and spiracle. Metasoma posteriad petiole pyriform, unsculptured, with row of setae evenly spaced on posterior margin of tergites; T2 spiracle laterally displaced. Ovipositor nearly straight, without dorsal node; ovipositor sheath setose throughout.

Hosts.

Agromyzidae . See comments under Eurytenes abnormis , the only species with host records.

Comments.

Fischer (1972) provides the most recent detailed description of Eurytenes s. str. (in German); ( Wharton (1988, 2006), Fischer (1998), and Wu and Chen (2006) provide diagnoses for Eurytenes s. str. and s. l. Wu and Chen (2006) were the first to use morphological features other than color for discriminating between species of Eurytenes s. str.

Key to New World species of Eurytenes sensu stricto

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1	Hind femur almost entirely dark brown ( Fig. 5 ); petiole entirely dark brown to black, more than twice longer than apical width. Mexico	Eurytenes ormenus
-	Hind femur yellow, sometimes slightly infuscated distally; petiole variable	2
2	Gena broad ( Fig. 28 ); petiole uniformly dark brown to black ( Fig. 29 ). Mexico	Eurytenes pachycephalus
-	Gena narrow ( Figs 8 , 15 , 24 ); petiole partly ( Figs 3 , 17 ) to entirely ( Fig. 13 ) yellow	3
3	Petiole yellow ( Fig. 13 ); clypeus somewhat chevron shaped ( Fig. 10 )	Eurytenes abnormis
-	Petiole mostly dark brown to black dorsally ( Fig. 17 ), yellow laterally ( Fig. 3 ); clypeus truncate or nearly so ventrally, broader and more nearly semi-circular ( Figs 1 , 18 )	4
4	Antenna with 31-35 flagellomeres; clypeus obviously infuscate dorsally ( Fig. 18 )	Eurytenes dichromus
-	Antenna with 29-31 flagellomeres; clypeus barely infuscate dorsally ( Fig. 1 )	Eurytenes microsomus

Figures

Figures 30 - 31. 30 Eurytenes dichromus sp. n., mesoscutum, dorsal-lateral oblique, arrow = notaulus 31 Eurytenes abnormis, mesosoma, dorsal view, arrow = scutellar sulcus.

Figures 4 - 7. Eurytenes spp., habitus. 4 Eurytenes abnormis, female 5 Eurytenes ormenus sp. n., female 6 Eurytenes pachycephalus sp. n., male 7 Eurytenes dichromus sp. n., female.

Figures 26 - 29. Eurytenes pachycephalus sp. n. 26 Face 27 Mesosoma, lateral view, arrow = precoxal sulcus 28 Head, lateral view 29 Petiole, dorsal view.

Figures 8 - 14. Eurytenes abnormis. 8 Head, lateral view 9 Mesosoma, lateral view, arrow = precoxal sulcus 10 Face 11 Propodeum, posterior view 12 Left fore and hind wing 13 Petiole, dorsal view, arrow = dorsope 14 Petiole, lateral view.

Figures 15 - 21. Eurytenes dichromus sp. n. 15 Head, lateral view 16 Mesosoma, lateral view, arrow = precoxal sulcus 17 Petiole, dorsal view 18 Face 19 Propodeum, posterior view 20 Petiole, lateral view 21 Fore and hind wing.

Figures 22 - 25. Eurytenes ormenus sp. n. 22 Face 23 Mesosoma, lateral view, arrow = precoxal sulcus 24 Head, lateral view 25 Propodeum, dorsal view, arrow = sculptured posterior-lateral field.

Figures 1 - 3. 1 Eurytenes microsomus sp. n., face, bars = clypeal measurements 2 Eurytenes dichromus sp. n., petiole, lateral view, bar = length measurement 3 Eurytenes microsomus, lateral view, arrow = precoxal sulcus, bar = mesosomal length.