Leiorhagium neteae, Haase, Martin & Zielske, Susan, 2015

Haase, Martin & Zielske, Susan, 2015, Five new cryptic freshwater gastropod species from New Caledonia (Caenogastropoda, Truncatelloidea, Tateidae), ZooKeys 523, pp. 63-87 : 71-73

Taxon classification Animalia Littorinimorpha Tateidae

Leiorhagium neteae sp. n.

Type material.

Holotype MNHN IM 2000-27867; paratypes MNHN IM 2000-27868 (20).

Type locality.

NeCa 44, Poya: stream at side of small road branching off road between Nétéa and Goipin toward the Vallée d’Adio, under shrub close to overgrown garden, 21°14'47.9"S, 165°15'45.0"E, 28 May 2012.

Etymology.

The new species is named after the village of Nétéa, which is closely proximal to our collecting locality.

Diagnosis.

Leiorhagium neteae sp. n. is very similar to Leiorhagium douii and Leiorhagium montfaouense . In Leiorhagium neteae sp. n. the palatal denticle is slightly larger and 1/8 whorl further behind the outher lip. The operculum has only a single denticle compared to 2-3 in Leiorhagium douii and Leiorhagium montfaouense . The distal loop of the renal oviduct of the new species forms a 270° loop counter-clockwise, whereas in the other two species this part of the oviduct is bent 180 ° clockwise. The penis of Leiorhagium neteae sp. n. is long and slender in contrast to the other species, where it has a broad base and a very long filament.

Shell.

Elongate-pupiform, 1.88 times higher than wide, 4.25-5 whorls, without colour, transparent; protoconch faintly pitted with c. 1 whorl; palatal denticle an elongate droplet c. 1/4 whorl behind outer lip; aperture slightly wider than high (Figs 2E, 3H, I, 4I, J).

Operculum.

Elongate-ellipsoidal, paucisprial, nucleus submarginal, orange, one non-calcareous white peg (N=4) (Fig. 5G, H).

External features.

Epidermis without pigment, eyes black.

Mantle cavity.

Ctenidium with 15 (1 male) or 19-22 (5 females) filaments; osphradium short-elongate, behind middle of ctenidium.

Digestive system.

Radula formula (N=4) (Fig. 6D): R: 4 1 4/2-3 2-3, L: 4-5 1 5, M1: 20-25, M2: 24-27; stomach without caecum; rectum close to pallial oviduct in females, with short loop left of prostate in male.

Female genitalia.

Ovary without lobes, proximal end 1.25-1.5 whorls below apex, comprising 0.25-0.5 whorls, not reaching stomach; anterior capsule gland yellow-orange, posterior capsule gland opaque-white, albumen gland milky-white; proximal loop of renal oviduct bent forward, distal loop short; bursa copulatrix globular, reaching slightly behind albumen gland; bursal duct long, entering anterior; no seminal receptacle (Fig. 7D).

Male genitalia.

Proximal end of lobate testis 1 whorl below apex, comprising slightly more than 0.5 whorls, covering proximal end of stomach; vesicula seminalis arising approximately in middle of testis; penis very long and slender (N=1) (Fig. 8E).

Remarks.

This is Leiorhagium sp. n. 5 of Zielske and Haase (2015). Leiorhagium neteae sp. n. is only known from the type locality.