Semiclaviger sikorae Wasmann, 1893

Hlaváč, Peter, Baňař, Petr & Parker, Joseph, 2013, The Pselaphinae of Madagascar. II. Redescription of the genus Semiclaviger Wasmann, 1893 (Coleoptera: Staphylinidae: Pselaphinae: Clavigeritae) and synonymy of the subtribe Radamina Jeannel, 1954, Zootaxa 3736 (3), pp. 265-276 : 271

Semiclaviger sikorae Wasmann, 1893

( Figs. 1–27 )

Semiclaviger sikorae Wasmann, 1893: 103

Semiclaviger sikorae Wasmann : Jeannel, 1954: 332; Jeannel, 1960: 209

Material studied: LECTOTYPE, 1 ♂, present designation: (h) Madagascar (Sikora) / (h) b. Cremastogaster Schenki Fr. / (p) red label TYPE / (h) Semiclaviger Sikorae Wasm , Type / (p) blue label MUSÉUM PARIS, 1952, COLL. R. OBERTHÜR / (p) red label Lectotype Semiclaviger sikorae Wasmann, P. Hlaváč & P. Baňař det., 2013. MNHN. PARALECTOTYPE, 1 ♂: (h) Madagasc. (Sikora) / (p) red label TYPE / (p) blue label MUSÉUM PARIS, 1952, COLL. R. OBERTHÜR / (p) red label Paralectotype Semiclaviger sikorae Wasmann, P. Hlaváč & P. Baňař det., 2013. MNHN. Other material: 3 ♂♂, 1 ♀: ABT/ Sept.2011/04 MADAGASCAR, AMBOHITANTELY Spec. Res., S18°11′46.4′′ E47°17′09.9′′, 1570m, 1.ix.2011 / sifting forest litter under Pandanus, Winkler app. extraction / L.S. Rahanitriniaina lgt. / Semiclaviger sikorae Wasmann, P. Hlaváč & P. Baňař det., 2013 (PHPC, MMBC); 1 ♀: ABT/ Sept.2011/12 MADAGASCAR, AMBOHITANTELY Spec. Res.; 3.ix.2011, S18°10′ 58.2″ E47°17′ 22.9″, 1522m / sifting forest litter by river, Winkler app. extraction, L.S. Rahanitriniaina lgt. / Semiclaviger sikorae Wasmann, P. Hlaváč & P. Baňař det., 2013 (MMBC), specimen gold-plated for SEM study;; 3 ♂♂, 1 ♀: ABT/ Nov.2011/11 MADAGASCAR, AMBOHITANTELY Spec. Res., 1497m, S18°10′52.6′′ E47°17′22.5′′, 18.xi.2011 / sifting forest litter, Winkler app. extraction, L.S. Rahanitriniaina lgt. / Semiclaviger sikorae Wasmann, P. Hlaváč & P. Baňař det., 2013 (PHPC, MMBC); 1 ♂: ABT/ Nov.2011/18 MADAGASCAR, AMBOHITANTELY Spec. Res., 1614m, S18°11′ 48.7″ E47°17′ 11.3″, 22.xi.2011 / sifting litter under group of palm trees, Winkler app. extraction, L.S. Rahanitriniaina lgt. / Semiclaviger sikorae Wasmann, P. Hlaváč & P. Baňař det., 2013 (MMBC), specimen gold-plated for SEM study.

Lectotype designation: Redescription of this species below is based on two male specimens deposited in MNHN having syntype status. Wasmann (1893) mentioned four specimens (3 males, 1 female) in his original description from the following locality: Andrangoloaka, O.-S-O. von Annanarivo, Madagascar. The locality labels of both specimens from Paris bear the following hand written text:? Madagascar (Sikora)’, lacking Andrangoloaka. Wasmann commonly provided type localities that were only communicated to him verbally. Because both specimens were collected by Sikora, we are of the opinion that they can be treated as syntypes. One male is here designated lectotype, the second male is paralectotype, in order to ensure the stability of the nomenclature and to provide a unique name-bearing type for Semiclaviger sikorae .

Description. Body length 1.20–1.30 mm, maximum width in posterior corners of elytra, about 0.75 mm. Long setae arranged in lines on pronotum and lateral part of elytra, elytral disc almost lacking setae. Visible part of head (in dorsal view) six times as wide as long, Pronotum slightly more than twice as wide as long and five times as long as head. Elytra about 1.90 times as wide as long and 1.35 times as long as pronotum. Abdomen about as long as elytra, only composite tergal plate visible dorsally, median distance between prebasal tergal keels slightly more than twice the distance of keel to lateral margin of paratergite.

Sexual dimorphism. Female with simple mesotibiae, lacking long preapical spur.

Host ant: Crematogaster (Crematogaster) schencki Forel, 1891

Bionomics. All newly collected specimens were sifted from leaf litter in primary forest in Ambohitantely Special Reserve in central Madagscar, with subsequent extraction via a Winkler apparatus.

Distribution. The species was described based on four specimens from Andrangoloaka [Andrangolaoka], O.-S.- O. [west-south-west] von Annanarivo [Antananarivo] ( Madagascar). The second locality is Gorge de la Mandraka, 1000 m (Jeannel, 1954: 332). The new record is from Ambohitantely Special Reserve, just to the north of Antananarivo. So far all known localities are in central Madagascar, close to Antananarivo, on the Central Plateau of the island ( Fig. 28 ).

Figures

FIGURE 1. Semiclaviger sikorae Wasmann, 1893, male, total dorsal view. Scale bar 1.0 mm.

FIGURES 2 – 7. Semiclaviger sikorae Wasmann, 1893. 2 – 3 male; 4 – 7 female. 2 — dorsal habitus; 3 — ventral habitus (III-IX — abdominal segments III-IX); 4 — head, anterior view (bc— bucal cavity, cl— clypeus, fr— frons); 5 — head, ventro-lateral view (cl— clypeus, gen— gena, gp— gular plate, occ— occipital constriction); 6 — trophi, antero-ventral view (bst— basistipes, cdcardo, lb— labrum, md— mandibula, mn— mentum, plp— palpomere, tp — tentorial pit); 7 — basis of antennae, anterior view (pd— pedicel, sc— scape).

FIGURES 8 – 15. Semiclaviger sikorae Wasmann, 1893. 8, 14 female; 9 – 13, 15 male. 8 — head, antero-dorsal view; 9 — dorsal face of head, dorso-lateral view; 10 — right antenna, dorsal view; 11 — head and pronotum, dorsal view; 12 — elytra, dorsal view; 13 — apex of left elytron with trichome, dorsal view; 14 — meso and metaventrite (cox II— mesocoxa, cox III — metacoxa, ististhmus, mtvp— metaventral process, tr II — mesotrochanter, tr III— metatrochanter); 15 — abdomen, dorsal view.

FIGURES 16 – 21. Semiclaviger sikorae Wasmann, 1893. 16 – 17, 21 female; 18 – 20 male. 16 — abdomen, ventral view; 17 — detail of surface of abdomen, ventral view; 18 — right fore tarsus, ventral view; 19 — basis of right fore tarsus, ventral view; 20 — right middle tibia, postero-dorsal view; 21 — right middle tarsus, anterior view.

FIGURES 22 – 24. Semiclaviger sikorae Wasmann, 1893, female. 22 — prosternum (bs— basisternal part, hy— hypomera, pccprocoxal cavity, pcs— procoxal socket, psc— prosternal carina); 23 — mesonotum (msc— mesoscutum, msl— mesoscutellum); 24 – abdomen, ventral view (blf— basolateral fovea, III-VI — sternites); 25 – terminalia, sternites VIII (VIII) and IX (IX) (pygidium) and tergite VIII (tg VIII). Scale bar in mm.

FIGURES 26 – 27. Semiclaviger sikorae Wasmann, 1893, male, aedeagus. 25 — dorsal view; 26 — lateral view. Scale bar in mm.

FIGURE 28. Distribution of Semiclaviger sikorae Wasmann, 1893 in Madagascar.