Engystomops pustulosus (Cope 1864)

Barrio-Amorós, César L., Rojas-Runjaic, Fernando J. M. & Señaris, J. Celsa, 2019, Catalogue of the amphibians of Venezuela: Illustrated and annotated species list, distribution, and conservation, Amphibian & Reptile Conservation (e 180) 13 (1), pp. 1-198 : 99

Engystomops pustulosus (Cope 1864)

Holotype: USNM 4339 (lost).

Type locality: “New Grenada, on the River Truando,” Colombia.

Distribution: Regions 1, 2, 3, 4, 5, 6. From Mexico through Central America to Colombia, Venezuela, Trinidad and Tobago, and Guyana in northern South America. Widespread in lowlands in northern Venezuela.

Remarks: May hide a complex of species (Wynn and Heyer 2002). At least two clear clades distinguishable, one from Southern Mexico to Costa Rica, another from Panama to northern South America (Weigt et al. 2005). Call has been studied in many localities ( Ryan et al. 2007).

Selected references: Lutz (1927); Ginés (1959); Rivero (1961, 1963a, 1964a,c,d); Heatwole et al. (1965); Tello (1968); Rivero and Esteves (1969); Lynch (1970); Staton and Dixon (1977); Hoogmoed and Gorzula (1979); Cannatella and Duellman (1984); Yústiz (1996); Duellman (1997); Gorzula and Señaris (1998); Nascimento et al. (2005); Weigt et al. (2005); Ryan et al. (2007); Tárano (2010).