Platypalpus anomalus, Barták & Kubík, 2018

Barták, Miroslav & Kubík, Štěpán, 2018, Hybotidae (Diptera) from Turkey, with descriptions of seven new species, Zootaxa 4410 (3), pp. 453-482: 456

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Platypalpus anomalus

sp. nov.

Platypalpus anomalus   sp. nov.

( Figs 5–7 View FIGURES 5–7 , 28 View FIGURES 27–30 )

Type material. HOLOTYPE ♂, labelled: “ Turkey: 8 km S of Çine , river bank, 68 m, 37°32′34′′N, 28°03′46′′E, 29.iv.–1.v.2016, Barták, Kubík” ( CULSP) GoogleMaps   . PARATYPES: 3♂, same data as holotype ( CULSP) GoogleMaps   .

Diagnosis. Small black species of the P. pallidiventris   - cursitans   group with a single pair of inclinate vertical setae; antenna with basal segments reddish yellow and the rest black, stylus greatly thickened; mesoscutum rather sparsely microtrichose; mid tibial spur short, short pale posteroventral setae on mid femur.

Description. Male. Head black, grey microtrichose. Frons about 0.05 mm broad and almost parallel-sided, widening slightly dorsally and 0.06 mm broad at level of anterior ocellus. Face 0.03 mm broad at middle. Gena very narrow and lustrous. Antenna with both basal segments reddish yellow, postpedicel and stylus black, postpedicel only slightly broader than basal part of stylus, 2.6–2.7X longer than wide, stylus greatly thickened (as in P. farabiensis   —see Shamshev 1998, fig. 7) and 1.4–1.5X longer than postpedicel. Clypeus microtrichose on anterior (dorsal) half and lustrous below (posteriorly). Palpus brown, ovate, half as long as labrum, with 2 long subterminal brownish yellow setae. Ocellar setae pale, about 0.09 mm long, second pair very short. Single pair of inclinate short vertical setae as long as ocellar, inserted wide apart (0.17 mm). Occiput short and sparsely brownish yellow setose dorsally and short paler setose ventrally. Proboscis black, half as long as head height. Thorax black, grey microtrichose, katepisternum narrowly lustrous, leaving microtrichose stripes both anteriorly and posteriorly. Mesoscutum sparsely covered with relatively long and adpressed microtrichia causing somewhat subshiny appearance, pleura more densely microtrichose. All thoracic setae pale, brownish yellow to yellow. Chaetotaxy: postpronotal seta long, additional setae very short; acrostichals narrowly biserial and diverging, short (0.06 mm), only about 5–6 setae in one row; dorsocentrals uniserial, evenly short and sparse anteriorly, posteriorly 1–2 longer pairs; notopleuron with 1–2 setae on posterior part, otherwise with only a few short setulae; 1 postalar and 1 pair of scutellar setae (with additional pair of small setulae laterally). Wing clear with yellowish veins, R4+5 and M1+ 2 in apical third almost parallel. Crossveins very broadly separated (on distance about 3– 4X longer than length of r-m crossvein (rather narrow second basal cell much longer than first basal cell), vein CuA1 vanishing before wing margin. CuA2 slightly recurrent and slightly C-shaped, anal vein indistinct in basal part. Costal seta brownish yellow, short. Squama whitish with yellow fringes. Halter whitish. Legs yellow including tarsi. Coxae yellow with yellow setae. Fore femur distinctly thickened in basal third, ventral setae short. Fore tibia slightly spindle-shaped dilated, with brownish setulae dorsally and anteriorly. Mid femur distinctly narrower than fore femur, with 6–8 short pale posteroventral setae, anteroventrally with several pale setae somewhat longer than those forming two rows of dark ventral spines. Mid tibia with short blunt-tipped apical spur and usual row of black spines ventrally. Hind legs thin, without conspicuous setae. Abdomen brown, lustrous except finely microtrichose sides of tergites 1–3. All abdominal setae pale (brown on genital lamellae) and short (except long marginal setae on last sternite). Genitalia ( Figs 5–7 View FIGURES 5–7 ): left epandrial lamella ( Fig. 7 View FIGURES 5–7 ) with long setae in dorsoapical part; cerci ( Fig. 6 View FIGURES 5–7 ) short and rather broad; right epandrial lamella ( Fig. 5 View FIGURES 5–7 ) with short surstylus. Female. Unknown. Length: body 2.0– 2.3 mm, wing 1.7–2.0 mm.

Etymology. The species name is derived from “ anomalus   ” (= anomalos, Greek adjective meaning anomalous, irregular) referring to several anomalous features of this species (shortened vein CuA1, greatly thickened stylus).

Distribution. Turkey, Aydın province.

Remarks. The species described above is closely allied to the Central Asian species, P. farabiensis Shamshev, 1998   , and P. tumidiarista Barták & Shamshev, 2015   . With darkened palpi, shape of left epandrial lamella, and narrower mid femur it is closer to the latter. From both species it differs in the structure of male genitalia (left epandrial lamella bears very long, dark posteriorly oriented setae in dorsal excision, broad apical projection, and much longer setae ventrally). Moreover, the lustrous spot on the katepisternum is broader than in both species with which it is compared (broader than posterior microtrichose stripe), vein CuA1, ending far before the wing margin (distance longer than lower crossvein), and clypeus is lustrous on ventral half. Platypalpus anomalus   sp. nov. leads to couplet 158 in the key by Grootaert & Chvála (1992), but differs from any other species of this section by its thickened stylus.