Amphibolips
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publication ID |
https://doi.org/ 10.11646/zootaxa.4877.1.1 |
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publication LSID |
lsid:zoobank.org:pub:84A56D01-FFCC-4D3B-95D8-36DA7AFEC0C7 |
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DOI |
https://doi.org/10.5281/zenodo.4567090 |
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persistent identifier |
https://treatment.plazi.org/id/467987FB-D44E-FFE8-ADD6-EF2238BDD0CD |
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treatment provided by |
Plazi |
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scientific name |
Amphibolips |
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Key to the Amphibolips View in CoL View at ENA adults of the ‘ nassa’ complex of Mexico and Central America
1. Metatarsal claw simple ( Fig. 8D View FIGURE 8 ). Antenna entirely light brown ( Fig. 8B View FIGURE 8 ) Mesopleuron weakly sculptured, speculum glabrous, smooth ( Fig. 7C View FIGURE 7 ). Mesoscutellum not depressed, or rarely slightly depressed posteriorly ( Fig. 7E View FIGURE 7 )..................... 2
1’. Metatarsal claw with large basal lobe ( Fig. 8E View FIGURE 8 ). Antennae dark at least on basal half, except in A. kinseyi n. sp. (brown on basal half and ambarine distally). Mesopleuron coarsely sculptured ( Fig. 9C View FIGURE 9 ). Speculum sculptured and sometimes pubescent. Me-soscutellum depressed posteriorly........................................................................ 3
2. Mesoscutum uniformly rugate. Notauli traceable only in posterior third, furrow deep, broad, and smooth. Median propodeal area smooth and glabrous. Metasomal terga smooth, without micropunctures. Forewing almost hyaline. Males and females........................................................................................... A. salicifoliae
2’. Mesoscutum not uniformly rugate, with transversal interrupted faint carinae on anterior half, with cells on posterior half ( Fig. 7E View FIGURE 7 ). Notauli complete, shallow, hardly traceable under sculpture, not emarginate by carinae, with rugose base ( Fig. 7E View FIGURE 7 ). Median propodeal area faintly rugose. Metasomal tergum II with fine micropunctures on posterior 1/3 ( Fig 7F View FIGURE 7 ), sometimes not visible in dorsal view, and rarely completely absent in the sexual generation; subsequent terga micropunctured. Forewing hyaline, with only a heavily infuscate patch present in the basal area of radial cell ( Fig. 8A View FIGURE 8 ). Sexual and asexual generations..................................................................................... A. quercuspomiformis comb. nov.
3. Forewing hyaline, only with a heavily infuscate patch in the basal area of radial cell ( Fig. 13A View FIGURE 13 ). Notauli emarginate, delimited by internal carina in posterior half........................................................................ 4
3’. Forewing with different colouration, infuscation pattern covering a greater area than basal area of radial cell ( Figs 13 View FIGURE 13 B–H). Notauli not emarginate, nor delimited by internal carina ( Figs 11 View FIGURE 11 A–K)........................................... 5
4. Metasoma reddish brown; posterior 1/3 of T2 with a band of hardly traceable micropunctures. Frons coarsely reticulate-rugose. Notauli complete. Mesoscutellum only slightly emarginate posteriorly ( Fig. 12A View FIGURE 12 ). Speculum finely rugose, glabrous. Males and females...................................................................................... A. aliciae
4’. Metasoma ambarine ( Fig. 6A View FIGURE 6 ); posterior 1/3 of T2 with a band of distinct micropunctures. Frons alutaceous with fine rugae ( Figs 6 View FIGURE 6 B–C). Notauli incomplete, visible under sculpture in posterior half ( Fig. 6D View FIGURE 6 ). Mesoscutellum emarginate posteriorly and laterally by a strong carina, delimiting posterior emargination ( Figs 6 View FIGURE 6 D–E). Speculum uniformly sparsely pubescent, with transversal carinae. Only females............................................................. A. kinseyi n. sp.
5. Forewing infuscate at least in distal half with a transverse hyaline band on the apical third of radial cell, extended towards inferior margin of wings or shorter in females ( Figs 1A View FIGURE 1 , 3E View FIGURE 3 , 10A View FIGURE 10 , 13B, 13D View FIGURE 13 ). Males forewing completely infuscate with a clear patch at least in median part of first cubital cell, sometimes extended to radial cell and posteriorly, sometimes reaching posterior margin ( Figs 2F View FIGURE 2 , 13C View FIGURE 13 ).................................................................................. 6
5’. Forewing lightly infuscate with a conspicuous heavily infuscate band in the anterior margin, never with a transverse hyaline band in females ( Figs 9A View FIGURE 9 , 13 View FIGURE 13 E–G). Males, forewing completely infuscate with a distinct darker band in the anterior margin, sometimes with a clear patch in distal half of radial cell, never extending posteriorly to costal cells ( Fig. 13H View FIGURE 13 )........... 13
6. Forewing with a narrow and incomplete hyaline band, not reaching the inferior margin; 2r vein conspicuously prolonged in medial part into radial cell ( Figs 10A View FIGURE 10 , 13 View FIGURE 13 C–D). Body black. Posterior sides of mesoscutellum, aside of posteromedian impression, strongly emarginate, dorsally V-shaped, with sharp horn projection in lateral view ( Figs 10D View FIGURE 10 , 12B, 12 View FIGURE 12 G–H). Propodeal carinae differentiated, subparallel ( Figs 10D View FIGURE 10 , 12I View FIGURE 12 )............................................................ 7
6’. Forewing with a wide and complete clear band, reaching the inferior margin; some males with completely infuscate forewing, without cross-band, with a small clear patch in median part of first costal cell near the margin with radial cell instead; 2r vein not prolonged into radial cell ( Fig. 13B View FIGURE 13 ). Head and mesosoma black, metasoma rufous or ambarine at least in some part ( Figs 1G View FIGURE 1 , 2E View FIGURE 2 , 3F View FIGURE 3 ). Posterior part of mesoscutum impressed but dorsally not V-shaped, without sharp horn projection in lateral view. Propodeal carinae indistinct............................................................................. 8
7. Forewing with anal and basal half of discoidal cells hyaline ( Figs 10A View FIGURE 10 , 13D View FIGURE 13 ). Area aside of parapsidal lines rugose ( Fig. 10C View FIGURE 10 ). Only females.............................................................................. A. turulli n. sp.
7’. Forewing with discoidal and anal cells completely infuscate ( Fig.13C View FIGURE 13 ). Area aside of parapsidal lines coriaceous ( Fig. 12B View FIGURE 12 ). Galls spherical, interior of the gall spongy, soft. Males and females....................................... A. dampfi
8. Notauli absent ( Fig. 11B View FIGURE 11 ; Nieves-Aldrey et al 2012: Fig. 2E View FIGURE 2 ). Mesoscutellum moderately emarginate posteriorly; scutellar foveae rugose, indistinctly separate medially ( Fig. 11B View FIGURE 11 ). Only females................................ A. durangensis
8’. Notauli distinct posteriorly and medially, anteriorly less defined or inconspicuous under the rugose sculpture ( Figs 1E View FIGURE 1 , 3C View FIGURE 3 , 11A, 11C View FIGURE 11 ). Mesoscutellum weakly emarginate posteriorly; scutellar foveae carinated, smooth or rarely rugose in males, but always divided by a distinct median carina, which is only rarely incomplete ( Figs 1E View FIGURE 1 , 3C View FIGURE 3 , 11A View FIGURE 11 )............................. 9
9. Mesoscutellum subquadrate posteriorly; scutellar foveae smooth. Posterodorsal area of mesopleuron smooth. Metasoma dorsally black, ventrally rufo-piceous. Costal cell of forewing completely infuscate ( Medianero & Nieves-Aldrey 2010: Figs 7 View FIGURE 7 A–C). Males and females................................................................... A. castroviejoi
9’. Mesoscutellum bilobate posteriorly ( Figs 1E View FIGURE 1 , 3C View FIGURE 3 , 11A View FIGURE 11 ); scutellar foveae with transverse carinae. Posterodorsal area of mesopleuron strongly rugose. Metasoma rufous. Costal cell of forewing not infuscate ( Fig. 13B View FIGURE 13 ). Only females.............. 10
10. Scutellar posteromedian depression absent or shallow, never reaching posteroventral margin of mesoscutellum ( Figs 11C, 11L View FIGURE 11 ). Only females................................................................................... A. fusus
10’. Scutellar posteromedian depression distinct, reaching posteroventral margin of mesoscutellum ( Figs 1E View FIGURE 1 , 3C View FIGURE 3 , 11A View FIGURE 11 )....... 11
11. Notauli complete, traceable under sculpture, reaching anterior margin ( Fig. 3C View FIGURE 3 ); median sulcus absent. Mesoscutellum 1.1x as long as broad; median carina between scutellar foveae faint ( Fig. 3C View FIGURE 3 ). Posterolateral projections of mesoscutellum extending downwards forming a carina emarginate posterior depression ( Fig. 3D View FIGURE 3 ). Only females................... A. bromus n. sp.
11’ Notauli incomplete, lost in sculpture in anterior half of mesoscutum, median sulcus distinct in female ( Fig. 11A View FIGURE 11 ); in males notauli and median sulci almost inconspicuous, covered by coarse rugulose sculpture ( Fig. 2D View FIGURE 2 ). Mesoscutellum in females 0.8–0.9x as long as broad, median carina between scutellar foveae strong ( Fig. 1E View FIGURE 1 ); in males mesoscutellum 1.1x as long as broad, median carina between scutellar foveae strong, partially covered by rugae of the foveae ( Fig. 2C View FIGURE 2 ). Posterior projections of mesoscutellum horn-like not extending downwards ( Figs 1E View FIGURE 1 , 11A View FIGURE 11 ) in both sexes................................ 12
12. Scutellar posterolateral projections pointed, elongated upwards ( Fig. 1E View FIGURE 1 ). Clypeus, mandibles uniformly chestnut brown ( Fig. 1D View FIGURE 1 ). Females with F11 clearly divided by incomplete sulcus; F11 2.0x as long as broad behind the sulcus. Metasoma ambarine ( Figs 1A, 1G View FIGURE 1 ). Males and females............................................................ A. bassae n. sp.
12’. Scutellar posterolateral projections rounded, not pointed upwards ( Fig. 11A View FIGURE 11 ). Clypeus chestnut brown, mandibles rufous. F11 not divided, with faint incomplete sulcus; F11 as long as broad after the sulcus. Metasoma reddish-brown. Only females............................................................................................... A. cibriani
13. Forewing with costal, basal, first cubital cells and part of anal cell under basal cell uniformly infuscate ( Fig. 13F View FIGURE 13 ). Only females....................................................................................... A. oaxacae
13’. Costal and anal cells not infuscate or less infuscate than basal and radial cell ( Fig. 13E View FIGURE 13 ). Some males always with completely infuscate forewing, not hyaline in third cubital and discoidal cells.............................................. 14
14. Proximal part of radial cell heavily infuscate; R1, Rs and 2r veins not traceable through infuscate band or hardly visible ( Fig. 13G View FIGURE 13 ). Forewing surface completely infuscate. Only females................................................... 15
14’. Radial cell not heavily infuscate, sometimes infuscate band lightened at distal end of radial cell usually extending to anterior margin of 3rd cubital cell; R1, Rs and 2r visible in females ( Fig. 13E View FIGURE 13 ); surface of forewing faintly infuscate, more or less hyaline. In males, forewing completely and heavily infuscate with veins at base of radial cell inconspicuous............... 16
15. Scutellar foveae emarginate medially by a conspicuous carina with transversal carinae emerging laterally ( Fig. 11K View FIGURE 11 ). Metasomal tergum II shining, smooth in front of micropunctures band................................... A. zacatecaensis
15’. Scutellar foveae smooth and not separated medially or with faint incomplete median carina on anterior margin, transversal carinae absent or very faint ( Fig. 11G View FIGURE 11 ). Metasoma tergum II posteriorly coriaceous in front of micropunctures band, not reaching half-length of tergum............................................................................ A. nassa
16. Scutellar foveae smooth, divided medially by a strong carina ( Fig. 11F View FIGURE 11 ). Scutellar posterior depression not reaching the basis of mesoscutellun, interrupted by strong transverse carina on posterior side ( Figs 5D View FIGURE 5 , 11F View FIGURE 11 ). Radial cell in males less than 3.5x as long as broad. Males and females............................................................ A. michoacaensis
16’. Scutellar foveae with transverse carinae extending laterally from median carina ( Figs 9F View FIGURE 9 , 11 View FIGURE 11 D–E, 11H–J). Scutellar depression reaching posterior margin of mesoscutellum, not interrupted by strong transverse carina ( Fig. 5C View FIGURE 5 ). Radial cell in males longer .................................................................................................. 17
17. Lateral margin of compound eye shining, smooth, delimited by conspicuous carina visible in anterior view ( Figs 9 View FIGURE 9 B–C)... 18
17’. Lateral margin of compound eye with fine sculpture not delimited by a carina or carina present only in the dorsal part, never visible in anterior view ( Figs 4 View FIGURE 4 A–B)..................................................................... 20
18. Metasomal tergum II punctured on posterior 1/5 on dorsal part and 1/3 laterally; with fine longitudinal striations on anterior part after punctuation. In females F1 1.4x as long as F2. Males and females................................. A. jaliscensis
18’. Metasomal tergum II forming a uniformly punctured band on posterior 1/3. In females F1 1.6x as long as F2. In some species males known also.................................................................................... 19
19. Metasomal tergum II with small dorsal projection of punctures towards anterior margin, never reaching half-length of tergum ( Fig. 9G View FIGURE 9 ); rest of tergum mainly smooth with fine coriaceous sculpture limiting punctures ( Figs 9E, G View FIGURE 9 ). Only females................................................................................................ A. rulli n. sp.
19’. Metasomal tergum II with band of punctures not projected on dorsal part; rest of tergum coriaceous reaching anterior margin of metasoma. Males and females................................................................. A. nevadensis
20. Mesoscutum rugose, posteromedial area less densely rugose, with larger cells compared to the rest of mesoscutum, usually along median carina and notauli ( Fig. 11J View FIGURE 11 ). Lateral and dorsal margin of eye with fine carinae irradiating from eye ( Fig. 4B View FIGURE 4 ). Scutellar foveae subtriangular, limited posteriorly and medially by rugose sculpture, medial carina shorter than length of foveae; posterior depression variable in depth, but never reaching scutellar foveae, uniformly rugulose with alutaceous sculpture between rugae like rest of mesoscutellum ( Figs 4D View FIGURE 4 , 11J View FIGURE 11 ). Metascutellum rectangular, more than 2.0x as broad as high ( Fig. 4F View FIGURE 4 ). Males and females............................................................................. A. tarasco
20’. Mesoscutum uniformly rugose, with similar cells, notauli and medial sulci inconspicuous ( Fig. 11D View FIGURE 11 ). Lateral margin of eyes rugose-coriaceous, not emarginate ( Fig. 4A View FIGURE 4 ). Scutellar foveae quadrangular, limited posteriorly by transverse carinae never by rugose sculpture, median carina as long as length of foveae; posterior depression reaching scutellar foveae, with transverse carinae, smooth interspaces ( Figs 11D View FIGURE 11 , 4C View FIGURE 4 ). Metascutellum quadrate, less than 1.8x as broad as high ( Fig. 4E View FIGURE 4 ). Only females........................................................................................ A. hidalgoensis
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