Quernocoris, WEIRAUCH, 2006

WEIRAUCH, CHRISTIAN E., 2006, New Genera, New Species, and New Combinations in Western Nearctic Phylini (Heteroptera: Miridae: Phylinae), American Museum Novitates 3521 (1), pp. 1-44 : 19-21

publication ID	https://doi.org/ 10.1206/0003-0082(2006)3521[1:NGNSAN]2.0.CO;2
persistent identifier	https://treatment.plazi.org/id/470287A9-FFED-FFAE-7171-FE2FC7B6AE42
treatment provided by	Carolina
scientific name	Quernocoris
status	gen. nov.

Treatment

Quernocoris View in CoL View at ENA , new genus

Figures 1 View Fig , 2 View Fig , 5–9 View Fig View Fig View Fig View Fig View Fig , 12 View Fig

TYPE SPECIES: Quernocoris caliginosus , new species.

DIAGNOSIS: Recognized by the moderate size, reddish to chocolate-brown coloration in the male (fig. 1), the pronotum mottled with dark spots, and the male genitalia with vesica J-shaped (fig. 5), with one apical blade, secondary gonopore of medium size, elongate ovoid and facing caudad, gonopore sclerite poorly defined. Distinguished from the similarly sized and colored Insulaphylus cruz (fig. 1) by the restriction of dark spots to the pronotum, which do not extend to the scutellum, and by the male genitalia.

DESCRIPTION: Male: Moderate size, ovoid and slightly elongate, rather stout in lateral view. COLORATION (fig. 1): General coloration reddish brown to dark brown, with anterior crescent-shaped portion of cuneus hyaline, remaining portion red. Head: Pale, with vertex with 4–5 dark transverse fasciae and one additional mark on interior margin of eye, distal part of clypeus and lateral longitudinal stripes brown, labrum light brown, mandibular plate orange, maxillary plate brown to dark brown suffused with red, gena and gula light brown to dark reddish brown, buccula white; antennal segment 1 yellowish with proximal and apical dark brown ring, segment 2 yellowish, suffused with brown at base and toward apex, segments 3 and 4 brownish; labial segments 1 to 3 yellow, segment 4 suffused with brown. Thorax: Pronotum, mesoscutum, and scutellum usually dark reddish brown, sometimes rather light orange brown, anterior pronotal lobe and to a lesser degree posterior lobe with irregular dark marks, mesoscutum with lateral margins orange, pleura in light specimens, propleura, dorsal half of mesepisternum, entire mesepimeron and metapleura yellowish suffused with red, with margins of procoxal cavity, dorsal margin of the mesepisternum, and posterior and dorsal portion of mesepimeron whitish, ventral half of mesepimeron brown, in dark specimens, pleura brown, with whitish on margin of procoxal cavity, dorsal margin of mesepisternum, posterior margin of mesepimeron, and area of evaporatory area, Legs: Legs yellowish or light brown, with brown spots on femora, distal portion of metafemur suffused with brown, base of tibiae with dark spot, bases of tibial spines with dark spots, distal tarsomeres suffused with brown, light specimens with bases of coxae suffused with brown, dark specimens with entire coxae brown. Hemelytra: Corium including clavus light reddish brown to dark chocolate brown, anterior rim of corium usually lighter than the remaining parts, cuneus with anterior crescent-shaped hyaline area, remaining portion bright red. Abdomen: Venter brownish suffused with red, pygophore yellow or light brown with dark marks. SURFACE AND VESTITURE (fig. 9E): Dorsum weakly shining, with rather dense, stout, subadpressed setae of moderate length, appearing either golden or black depending on angle of observation, vertex and pronotum also with sparse suberect, stout, black setae. STRUCTURE: Head (figs. 2E, 5): Head triangular in dorsal aspect, vertex wide, slightly convex and almost straight behind, clypeus weakly produced, mandibular plate slightly produced, maxillary plate sunken, buccula short, buccal cavity large and ovoid, gula short; eyes about 3/4 as high as head; large, weakly emarginate posterior of antennal fossa, posterolateral margins contiguous with anterolateral margins of pronotum; antennal segment 1 short and relatively slender, segment 2 long, slender at base and with increasing diameter toward apex, segments 3 and 4 together about 3/4 of length of segment 2; apex of labium reaching base of metacoxa. Thorax (figs. 7E, 8E): Pronotum trapeziform, anterior margin straight, lateral margins slightly convex, posterior margin slightly sinuate, anterior and posterior pronotal lobes not demarcated, callus not developed, metapleural evaporatorium with area of mushroomlike cuticle roughly triangular, mushroomlike cuticle anterior to mesothoracic spiracle consisting of few, rather large elevations (fig. 7E). Legs: Slender; claws slender, pulvilli of moderate size, covering about half of ventral claw surface, parempodia setiform, slender and moderately elongate (fig. 8E). Hemelytra: Hemelytra subparallel, cuneus elongate triangular. Abdomen: Abdomen reaching apex of cuneus. GENITALIA (fig. 5): Pygophore: Pygophore rather stout and large. Parameres: Right paramere broad and lanceolate; left paramere with anterior arm rather short with pointed apex, posterior arm moderately long, tapering and slightly bent ventrad. Phallotheca: Moderately slender and elongate, almost straight, without apparent ornamentation, ventral opening slitlike. Vesica: Vesica J-shaped, with one apical blade, secondary gonopore situated in apical quarter, of medium size, elongate ovoid and facing caudad, gonopore sclerite poorly defined.

Female (fig. 1): General coloration reddish, much lighter than male. GENITALIA (fig. 6): See description of type species Quernocoris caliginosus , new species.

ETYMOLOGY: Named for its occurrence on oaks, after the Latin adjective quernus, meaning ‘‘of oak’’. The gender of the name is masculine.

DISCUSSION: The monotypic genus Quernocoris is rather widespread in mainland California, but also occurs on the Channel Islands. According to Powell (1994) this pattern of distribution is found in most insects recorded from the California Channel Islands. A similar distribution is also seen in two species of the Mirinae genus Irbisia Reuter ( Schwartz 1984) .

Quernocoris caliginosus , new species

Figures 1 View Fig , 2 View Fig , 5–9 View Fig View Fig View Fig View Fig View Fig , 12 View Fig

HOLOTYPE: Male: USA: California: Orange Co.: Cleveland National Forest, El Cariso Campground, 33.3 ° N 116.8 ° W, 750 m, 12 May 1978, J. D. Pinto and R. T. Schuh, Quercus sp. (Fagaceae) , 13 ( AMNH _PBI 00095227) ( AMNH).

DIAGNOSIS: As in generic diagnosis.

DESCRIPTION: Male: As in generic description, total length 3.21–3.88, length from apex of clypeus to cuneal fracture 2.25–2.73, width across pronotum 1.06–1.24. COLORATION (fig. 1): General coloration as in generic description. Head: Head including antennae and labium as in generic description. Thorax: Thorax including legs and hemelytra as in generic description. Abdomen: Venter as in generic diagnosis. SURFACE AND VESTI- TURE (fig. 9E): Dorsum and hemelytra as in generic description. STRUCTURE: Head (figs. 2E, 5): Head including eyes, antennae, and labium as in generic description. Thorax (figs. 7E, 8E): Thorax including hemelytra and legs as in generic description. Abdomen: Abdomen including genitalia as in generic description. GENITALIA (fig. 5): As in generic description.

FEMALE (fig. 1): Female as in generic description. Total length 3.31–3.59, length of apex clypeus-cuneal fracture 2.26–2.55, width across pronotum 1.15–1.19. GENITALIA (fig. 6): Vestibulum stout, twisted S-shaped, bursa copuatrix rather large, with posterior margin having distinct paramedian lobes, sclerotized rings medium size.

ETYMOLOGY: Named for the dark brown coloration of the male, after Latin adjective caliginosus , meaning ‘‘dark’’.

HOSTS: Recorded from various species of Quercus , among them Quercus agrifolia Née , Q. dumosa Nutt. , Q. engelmanni Greene , and Q. kellogii Newb.

DISTRIBUTION: Distributed in the Coast Ranges south of San Francisco, the mountainous regions in southern California, extending south to San Diego County, with records from Santa Cruz and Santa Catalina islands, but also two records from the foothills of the Sierra Nevada.

DISCUSSION: This species is sympatric with Insulaphylus cruz , on Santa Cruz Island, with both species recorded from the same host. In San Francisco, it is sympatric with Vesperocoris paddocki , new combination, where both species occur on Quercus agrifolia .

PARATYPES: USA: California: Los Angeles Co.: Turnbull Canyon, 33.98472 ° N 118.03139 ° W, 7 Apr 1936, E. L. Paddock, Quercus sp. (Fagaceae) , 43 (AMNH_PBI 00068810–AMNH_PBI 00068813), 4♀ (AMNH_PBI 00068814–AMNH_PBI 000- 68817) (USNM). Orange Co.: Cleveland National Forest, El Cariso Campground, 33.3 ° N 116.8 ° W, 750 m, 12 May 1978, J. D. Pinto and R. T. Schuh, Quercus sp. (Fagaceae) , 53 (AMNH_PBI 00058807–AMNH_PBI 00058811), 1♀ (AMNH_ PBI 00058813) (AMNH). Quercus sp. (Fagaceae) , 13 (AMNH_PBI 00058806), 1♀ (AMNH_PBI 00058812) (CNC). San Bernardino Co.: 4 mi E of Mentone, 34.07 ° N 117.0638 ° W, 750 m, 11 May 1978, J. D. Pinto, Quercus sp. (Fagaceae) , 83 (AMNH_PBI 00082363–AMNH_PBI 00082368, AMNH_PBI 00082374–AMNH_PBI 00082375), 10♀ (AMNH_PBI 00082376–AMNH_PBI 000- 82385) (UCR). San Diego Co.: just above Lake Henshaw on S7 (also known as Henshaw Dam), 33.24167 ° N 116.76222 ° W, 900 m, 30 Apr 1985, R. T. Schuh and B. M. Massie, Quercus dumosa (Fagaceae) , det. K. Nixon 1985, 13 (AMNH_PBI 00058788), 1♀ (AMNH_PBI 00058790) (AM). Quercus dumosa (Fagaceae) , det. K. Nixon 1985, 93 (AMNH_PBI 00058778–AMNH_PBI 000- 58786), 8♀ (AMNH_PBI 00058792–AMNH_PBI 00058799) Quercus dumosa (Fagaceae) , det. K. Nixon 1985, 93 (AMNH_PBI 00058778– AMNH_PBI 00058786), 8♀ (AMNH_PBI 000- 58792–AMNH_PBI 00058799) (AMNH). Quercus dumosa (Fagaceae) , det. K. Nixon 1985, 13 (AMNH_PBI 00058787), 1♀ (AMNH_PBI 000- 58791) (ZISP). Santa Barbara Co. : Santa Cruz Island, Central Valley, 34.01861 ° N 119.68083 ° W, 23 Apr 1976, J. D. Pinto, Quercus dumosa (Fagaceae) , 63 (AMNH_PBI 00082430–AMNH_ PBI 00082435), 2♀ (AMNH_PBI 00082436, AMNH_PBI 00082437) (UCR). Ventura Co.: Ojai, 34.44806 ° N 119.24194 ° W, 16 Apr 1969, R. Shaver and L. Paddock, Quercus kellogii (Fagaceae) , 43 (AMNH_PBI 00073794–AMNH_PBI 000- 73797), 4♀ (AMNH_PBI 00073798–AMNH_PBI 00073801) (CAFA).

OTHER SPECIMENS EXAMINED: USA: California: Alameda Co.: Berkeley, 37.87167 ° N 122.27167 ° W, 22 Apr 1915, E. P. Van Duzee, 13 (AMNH_PBI 00079344) (UCB). Inyo Co. : Independence, 36.80278 ° N 118.19917 ° W, 14 Jun 1929, R. L. Usinger, 13 (AMNH_PBI 00077569) (CAS). Kern Co.: Lebec, 34.84167 ° N 118.86389 ° W, 1219 m, 15 May 1928, J. O. Martin, 13 (AMNH_PBI 00077568) (CAS). S of Tehachapi on Water Canyon Rd, 35.09563 ° N 118.4909 ° W, 1445 m, 21 May 2004, Schuh, Cassis, Schwartz, Weirauch, Wyniger, Forero, Quercus john-tuckeri Nixon and Muller (Fagaceae) , det. K. Nixon 2004, 23 (AMNH_PBI 00095077, AMNH_PBI 000- 95078), 6♀ (AMNH_PBI 00095079–AMNH_PBI 00095084) (AMNH). Los Angeles Co.: Glendora, 34.13611 ° N 117.86444 ° W, 2 May 1937, B. E. Norland, 13 (AMNH_PBI 00074198) (LACM).

Text continues on page 33 Oaks, Mint Canyon, 34.41528 ° N 118.45278 ° W, 20 Apr 1932, E. P. Van Duzee, 23 (AMNH_PBI 00077573, AMNH_PBI 00077574), 2♀ (AMNH_ PBI 00077575, AMNH_PBI 00077576) (CAS). Pasadena, 34.14778 ° N 118.14361 ° W, 9 Apr 1909, Grinnell, 23 (AMNH_PBI 00077264, AMNH_PBI 00077265), 1♀ (AMNH_PBI 00077564) (CAS); 1 May 1909, Grinnell, 63 (AMNH_PBI 00077266– AMNH_PBI 00077268, AMNH_PBI 00077557– AMNH_PBI 00077559), 3♀ (AMNH_PBI 000- 77560–AMNH_PBI 00077562) (CAS); 5 Jun 1909, Grinnell, 33 (AMNH_PBI 00077269–AMNH_PBI 00077270, AMNH_PBI 00077563) (CAS); 25 May 1909, Grinnell, 13 (AMNH_PBI 00077271) (CAS). Santa Catalina Island, Avalon Canyon, 33.38333 ° N 118.41667 ° W, 31 Mar 1968, J. Powell, 23 (AMNH_PBI 00079287, AMNH_PBI 00079288), 2♀ (AMNH_PBI 00079290, AMNH_PBI 000- 79291) (UCB). Santa Catalina Island, Little Harbour, 33.38333 ° N 118.41667 ° W, 1 Apr 1968, P. A. Opler, 23 (AMNH_PBI 00079292, AMNH_ PBI 00079293) (UCB). Santa Catalina Island, Pebbly Beach Canyon, 33.38333 ° N 118.41667 ° W, 31 Mar 1968, P. A. Opler, catalina cherry, 13 (AMNH_PBI 00079289) (UCB). Saugus 5 mi E Mint Cn., 34.41139 ° N 118.53917 ° W, 20 Apr 1932, E. P. Van Duzee, 13 (AMNH_PBI 00077572), 2♀ (AMNH_PBI 00077577, AMNH_PBI 00077578) (CAS). Tanbark Flat, 33.69111 ° N 116.67056 ° W, 17 Jun 1958, A. S. Menke, 13 (AMNH_PBI 000- 77571) (CAS). Whittier, 33.97917 ° N 118.03194 ° W, 18 Apr 1935, E. L. Paddock, Quercus sp. (Fagaceae) , 13 (AMNH_PBI 00068818), 1♀ (AMNH_PBI 00068819) (USNM). Marin Co.: Fairfax, 37.98722 ° N 122.58778 ° W, 7 May 1911, E. C. Van Dyke, 1♀ (AMNH_PBI 00077565) (CAS). Terra Linda E of Rt 101 at Freitas Prwy, 38.00417 ° N 122.54861 ° W, 25 Apr 1980, Russell and Schwartz, 3♀ (AMNH_PBI 00058814–AMNH_ PBI 00058816) (AMNH). Monterey Co.: 2 miles NE Bryson, 35.82717 ° N 121.06414 ° W, 17 Apr 1966, C. W. O’Brien, 23 (AMNH_PBI 00079351, AMNH_PBI 00079352) (UCB). Bradley, 35.86333 ° N 120.79972 ° W, 23 Apr 1962, E. P. Van Duzee, 23 (AMNH_PBI 00077566, AMNH_PBI 00077567) (CAS). Wiley Ranch 6 mi W Greenfield, 36.32078 ° N 121.35078 ° W, 366 m, 3 May 1975, J. Powell, 1♀ (AMNH_PBI 00079282) Quercus douglasii (Fagaceae) , 83 (AMNH_PBI 00079272– AMNH_PBI 00079278, AMNH_PBI 00079618) (UCB); 2 May 1975, S. Szerlip, Quercus douglasii (Fagaceae) , 13 (AMNH_PBI 00079279), 2♀ (AMNH_PBI 00079280, AMNH_PBI 00079281) (UCB); 3 May 1975, W.W. Middlekauff, 13 (AMNH_PBI 00079353), 1♀ (AMNH_PBI 000- 79354) (UCB); 3 May 1975, [collector unknown], 23 (AMNH_PBI 00079356, AMNH_PBI 000- 79357) (UCB). Orange Co.: Cleveland National Forest, El Cariso Campground, 33.3 ° N 116.8 ° W, 750 m, 12 May 1978, J. D. Pinto and R. T. Schuh, Quercus sp. (Fagaceae) , 1♀ (AMNH_PBI 000- 95230) Quercus dumosa (Fagaceae) , det. K. Nixon 1985, 13 (AMNH_PBI 00095228) Quercus engelmanni (Fagaceae) , det. K. Nixon 1985, 1♀ (AMNH_PBI 00095231) (AMNH). Santa Ana Mountains , 33.75 ° N 117.54167 ° W, 29 Apr 1962, B. Ewing, 13 (AMNH_PBI 00082643) (UCR). Riverside Co.: 2 mi N Poppet Flat on Hwy 243, 33.87901 ° N 116.85167 ° W, 22 May 1976, J. D. Pinto, Quercus agrifolia (Fagaceae) , 1♀ (AMNH_ PBI 00082639) (UCR). 2 mi N of Poppet Flat on Rt 243, 33.87891 ° N 116.85167 ° W, 22 May 1976, J. D. Pinto, Quercus dumosa (Fagaceae) , 43 (AMNH_PBI 00082386–AMNH_PBI 00082389), 1♀ (AMNH_PBI 00082390) (UCR). 6 mi W of Murrieta, Santa Rosa Plateau Area, 33.55 ° N 117.32074 ° W, 625 m, 1 May 1985, R. T. Schuh and B. M. Massie, Quercus dumosa (Fagaceae) , det. K. Nixon 1985, 1♀ (AMNH_PBI 00059000) Quercus engelmanni (Fagaceae) , det. K. Nixon 1985, 23 (AMNH_PBI 00058998, AMNH_PBI 00058999), 24♀ (AMNH_PBI 00058822– AMNH_PBI 00058845) Quercus engelmanni Greene (Fagaceae) , det. K. Nixon 1985, 23 (AMNH_PBI 00058998, AMNH_PBI 000- 58999), 24♀ (AMNH_PBI 00058822–AMNH_ PBI 00058845) Quercus englemanni Greene (Fagaceae) , det. K. Nixon 1985, 13 (AMNH_PBI 00060493) (AMNH). Cleveland National Forest, El Cariso Campground on Hwy 74, 33.65889 ° N 117.40972 ° W, 762 m, 12 May 1978, J. D. Pinto, Quercus sp. (Fagaceae) , 23 (AMNH_ PBI 00082391, AMNH_PBI 00082392), 6♀ (AMNH_PBI 00082393–AMNH_PBI 00082398) (UCR). Poppet Flat, 33.85 ° N 116.85167 ° W, 22 May 1976, G. Clark, 13 (AMNH_PBI 00082429) (UCR); 22 May 1976, B. A. Bowers, 13 (AMNH_PBI 00082672), 20♀ (AMNH_PBI 00082652–AMNH_PBI 00082671) (UCR). San Jacinto Mountains, junction Poppet Flat Road and Route 243, 33.75028 ° N 116.66667 ° W, 20 May 2000, M. D. Schwartz, Quercus palmeri (Fagaceae) , det. L. Raz 2002, 63 (AMNH_PBI 00059002– AMNH_PBI 00059006, AMNH_PBI 00059008), 7♀ (AMNH_PBI 00059009–AMNH_PBI 000- 59015) Rhamnus tomentella (Rhamnaceae) , det. L. Raz 2002, 13 (AMNH_PBI 00059007) (AMNH). Santa Rosa Plateau Reserve , 33.52417 ° N 117.27528 ° W, 29 Apr 1985, J. D. Pinto, Quercus engelmanni (Fagaceae) , 23 (AMNH_PBI 000- 82677, AMNH_PBI 00082678), 1♀ (AMNH_PBI 00082679) (UCR). Sendie Creek, 1 May 1979, H. L. Murray, 13 (AMNH_PBI 00082642) (UCR). San Bernardino Co.: 4 mi E of Mentone, 34.07 ° N 117.0638 ° W, 750 m, 11 May 1978, J. D. Pinto, Quercus sp. (Fagaceae) , 13 (AMNH_PBI 00082686), 1♀ (AMNH_PBI 00082723) (UCR). San Diego Co.: 3.5 mi S of Lake Henshaw on S7 (also known as Henshaw Dam), 33.19107 ° N 116.76222 ° W, 1220 m, 30 Apr 1985, R. T. Schuh and B. M. Massie, Quercus engelmanni (Fagaceae) , det. K. Nixon 1985, 33 (AMNH_PBI 00058800– AMNH_PBI 00058801, AMNH_PBI 00095092), 3♀ (AMNH_PBI 00058802–AMNH_PBI 000- 58803, AMNH_PBI 00095229) Quercus engelmanni Greene (Fagaceae) , det. K. Nixon 1985, 33 (AMNH_PBI 00058800–AMNH_PBI 00058801, AMNH_PBI 00095092), 3♀ (AMNH_PBI 00058802–AMNH_PBI 00058803, AMNH_PBI 00095229) (AMNH). 7 mi S of Warner Springs, Carrizo Creek intersection w/ Rt 79, 33.16587 ° N 116.63707 ° W, 16 May 1982, M. D. Schwartz, Quercus engelmanni Greene (Fagaceae) , 33 (AMNH_PBI 00058846–AMNH_PBI 00058848), 3♀ (AMNH_PBI 00058849–AMNH_PBI 000- 58851) (AMNH). Palomar Mountain, 33.36333 ° N 116.83528 ° W, 28 Jun 1963, J. Powell, 33 (AMNH_PBI 00079341–AMNH_PBI 00079343) (UCB). San Diego County, 32.71528 ° N 117.15639 ° W, 6 May 1913, E. P. Van Duzee, 13 (AMNH_PBI 00077356) (CAS). Just above Lake Henshaw on S7 (also known as Henshaw Dam), 33.24167 ° N 116.76222 ° W, 900 m, 30 Apr 1985, R. T. Schuh and B. M. Massie, Quercus dumosa (Fagaceae) , det. K. Nixon 1985, 33 (AMNH_PBI 00058789, AMNH_PBI 00059001, AMNH_PBI 00095226) (AMNH). 13 (AMNH_PBI 00072153) (CNC). San Luis Obispo Co.: 12.3 mi E of Arroyo Grande, Huasna Valley, 35.11841 ° N 120.37158 ° W, 310 m, 9 May 1985, R. T. Schuh and B. M. Massie, 1♀ (AMNH_PBI 00058805) (AMNH). Arroyo Grande Creek SW of San Luis Obispo, 35.11644 ° N 120.58609 ° W, 160 m, 8 May 1985, R. T. Schuh and B. M. Massie, Quercus lobata (Fagaceae) , 13 (AMNH_PBI 00059055) (AMNH). Avenales Guard Station Jct, Huasna Valley E of Arroyo Grande, 35.11861 ° N 120.58972 ° W, 840 m, 9 May 1985, R. T. Schuh and B. M. Massie, 23 (AMNH_PBI 00059052, AMNH_PBI 00059053) Quercus douglasii H. and A. ( Fagaceae ), 23 (AMNH_PBI 00058817, AMNH_PBI 00058818) (AMNH). E of Arroyo Grande, Husana Valley, Stony Creek Campground, 35.20933 ° N 120.25846 ° W, 9 May 1985, R. T. Schuh and B. M. Massie, Quercus agrifolia oxyadenia (Fagaceae) , det. K. Nixon 1985, 13 (AMNH_PBI 00059054), 1♀ (AMNH_PBI 00059056) (AMNH). Poly. Campus, Poly Cyn, 38.00222 ° N 120.13611 ° W, 125 m, 8 May 1985, R. T. Schuh and B. M. Massie, 13 (AMNH_PBI 00058819) (AMNH). San Luis Obispo, 35.28278 ° N 120.65861 ° W, 24 Apr 1919, E. P. Van Duzee, 13 (AMNH_PBI 00077570) (CAS). Santa Barbara Co. : Los Prietos Ranger Station, 34.54222 ° N 119.78472 ° W, 14 Mar 1967, J. Powell, Quercus douglasii (Fagaceae) , 13 (AMNH_PBI 00079286) (UCB). Santa Cruz Island, 34.01667 ° N 119.71667 ° W, 23 Apr 1976 – 26 Apr 1976, J. D. Pinto, Quercus agrifolia (Fagaceae) , 13 (AMNH_PBI 00082637), 1♀ (AMNH_PBI 00082638) (UCR); 24 Mar 1941, G. P. Kanakoff, Quercus sp. (Fagaceae) , 2♀ (AMNH_PBI 00074199, AMNH_PBI 00074200) (LACM); 15 Apr 1973, M. L. May, 13 (AMNH_PBI 000- 82428) (UCR). Santa Cruz Island, Canada Cervada, 34.01667 ° N 119.71667 ° W, 26 Apr 1966, P. Rude, 13 (AMNH_PBI 00079581) (UCB). Santa Cruz Island, Cascada, 34.01667 ° N 119.71667 ° W, 1 May 2000, J. Powell, Light Trap, 13 (AMNH_PBI 00079340) (UCB). Santa Cruz Island, Central Valley, 34.01861 ° N 119.68083 ° W, 23 Apr 1976, J. D. Pinto, 13 (AMNH_PBI 00082438), 3♀ (AMNH_PBI 00082439–AMNH_ PBI 00082441) (UCR); 25 Apr 1966, J. Slater, Quercus sp. (Fagaceae) , 13 (AMNH_PBI 000- 79580) (UCB). Santa Cruz Island, Central Valley, 34.01667 ° N 119.71667 ° W, 29 Apr 1966, J. Powell, 33 (AMNH_PBI 00079346, AMNH_PBI 000- 79577, AMNH_PBI 00079617), 1♀ (AMNH_PBI 00079347) (UCB); 23 Apr 1976 – 26 Apr 1976, B. A. Bowers, 23 (AMNH_PBI 00082634, AMNH_PBI 00082635), 1♀ (AMNH_PBI 00082636) (UCR); 23 Apr 1976 – 26 Apr 1976, P. McNalley, 13 (AMNH_PBI 00082632), 1♀ (AMNH_PBI 00082633) (UCR); 25 Apr 1966, R. L. Langston, 1♀ (AMNH_PBI 00079619) (UCB). Santa Cruz Island, Field Station, 34.01667 ° N 119.71667 ° W, 23 Apr 1976 – 26 Apr 1976, G. Clark, 1♀ (AMNH_PBI 00082640) (UCR). Santa Cruz Island, Prisoner’s Harbor Cr, 34.01667 ° N 119.71667 ° W, 16 Mar 1969, J. Powell, 23 (AMNH_PBI 00079283, AMNH_PBI 00079284), 1♀ (AMNH_PBI 000- 79285) (UCB). Santa Cruz Island, Ridge N of Laguna Cyn., 34.01667 ° N 119.71667 ° W, 28 Apr 1966, J. Wolf, 13 (AMNH_PBI 00079579) (UCB); 28 Apr 1966, J. A. Slater, 1♀ (AMNH_PBI 00079348) (UCB); 28 Apr 1966, P. Rude, 13 (AMNH_PBI 00079578) (UCB); 28 Apr 1966, J. Powell, 1♀ (AMNH_PBI 00079583) (UCB). Santa Cruz Island, Upper Central Valley, 34.01861 ° N 119.68083 ° W, 305 m, 26 Apr 1966, J. Powell, 33 (AMNH_PBI 00079330–AMNH_PBI 00079332), 7♀ (AMNH_PBI 00079333–AMNH_PBI 000- 79339) (UCB); 26 Apr 1966, P. A. Rude, 23 (AMNH_PBI 00079349, AMNH_PBI 00079350), 1♀ (AMNH_PBI 00079584) (UCB). Upper Oso Campground off Rt 154, 34.55583 ° N 119.75389 ° W, 305 m, 7 May 1985, R. T. Schuh and B. M. Massie, Quercus agrifolia oxyadenia Torr. (Fagaceae) , det. K. Nixon 1985, 1♀ (AMNH_PBI 00058804) (AMNH). Santa Clara Co. : Stanford University Campus, 37.42722 ° N 122.16917 ° W, 29 Apr 1928, R. L. Usinger, 13 (AMNH_PBI 00079582) (UCB). Ventura Co.: Tule Creek, 34.55917 ° N 119.26849 ° W, 27 Jun 1965, P. M. Jump, 13 (AMNH_PBI 00074196) (LACM).