Cimeliomorpha perspinosa, Pinkaew & Horak, 2019

Pinkaew, Nantasak & Horak, Marianne, 2019, Revision of the genus Cimeliomorpha Diakonoff (Lepidoptera: Tortricidae), Zootaxa 4615 (3), pp. 457-480 : 475-477

Cimeliomorpha perspinosa , sp. n.

( Figs. 8 , 20−21 , 31−32, 39, 55−58)

Thorax. Without raised posterior scale tufts; pronotal collar, tegula and mesonotum yellow. Forewing broadly subtriangular, length 9.6 mm in male (n = 1) ( Fig. 20 ), 9.9 mm in female (n= 1) ( Fig. 21 ); costa evenly curved, male costal fold absent, termen rather round, weakly concave below apex, basal 2/5 of wing yellow, distal 3/5 of wing with orange brown ground colour, with a broad transverse silvery band along its inner margin, angled near costa, inwardly followed by broad orange brown band in male, but inner margin of transverse silvery band in female edged with a narrow black line; with five additional oblique silvery lines from costa, first ending in silvery spot at R 4, second narrow and short extending to R 2, third from 3/4 costa to termen at M 2, the fourth narrow and short extend- ing to R 4, fifth across apex preceded by a parallel yellow subtriangular mark; ocelloid patch a large roundish area of white-tipped orange brown scales producing fine transverse white lines, with fine longitudinal blackish striation, scattered with blackish dots, distally delineated by two convergent silvery streaks; with a smaller central and four larger spots of raised silvery scales, shadowed by black and some orange, an angled series of 3 behind CuA 1, fifth at the anterior angle of the discal cell, costa with 8 minute dark brown dots along basal 2/5, remainder with black spots alternating with yellow spots; fringe brown with dark basal line. Underside of forewing orange with dark brown around margin except basal half of dorsum, with small yellowish grey triangles alternating with black spots along costa. Hindwing orange with blackish band along edge except in anal area; wing margin edged with narrow orange line, inner margin of blackish band roughly rectangular; fringe brown, with dark basal line to CuP, then gradually paler to white. Underside of hindwing orange, blackish margin roughly as on upper surface extending along edge except along anal margin, costa with blackish subelliptical patch medially and weakly connected with black apex.

Abdomen. Male genitalia ( Figs. 31−32 , 39 ) with tegumen moderately sclerotised, short and wide, dorsally round with dense setae, pedunculus with moderately large anterior narrowly triangular process; uncus absent; socii moderately large, lobe-shaped, hairy; gnathos membranous arising from near middle of tegumen; vinculum a mod- erately long, curved band, moderately sclerotised; juxta rather small, caulis rather short; phallus slender, moderately long and curved, with ventrodistal point; anellus surrounding base of phallus and extending obliquely, membrane next to juxta with numerous short setae; valva with large basal opening and deep emargination at base of cucullus, sacculus large, subrectangular, basally with short, sparse setae, dorsal margin with dense scale sockets, and with group of moderately dense scale sockets on ventral margin of basal opening; cucullus rather small, with rounded apex, outer surface with dense scale sockets, inner surface with dense bristles. Female genitalia ( Figs. 55−58 ) with papillae anales densely setose; tergum VIII with moderately dense scale sockets posteriorly and on lateral triangular extensions; sterigma large, semicircular, strongly sclerotised, posterior margin lightly sinuate, with dense scale sockets except anteriorly and on raised rim dorsally around ostium; sternum VII sclerotised especially along anterior margin, posterior margin with deep, rectangular excavation, with very dense scale sockets medially and along posterior margin; colliculum long and slender, sclerotised; ductus bursae very narrow except for granulate anterior fourth; ductus seminalis arising from anterior 1/3 of ductus bursae; corpus bursae ovate, shorter than length of ductus bursae, with two nearly equal, sinuate, horn-shaped signa.

Holotype. ♂, [West Papua] New Guinea: Mimika R. VII 1910, F.R. Wollaston, 1911-229, 350405, B.M. geni- talia slide no. 33123 ( NHMUK).

Other material examined. Papua New Guinea: 1♀, Bastian [New Britain], Doherty, 1892, no. 42356, Wals- ingham Collection 1910-427, B.M. genitalia slide no. 33122 ( NHMUK) .

Etymology. The specific epithet refers to the cluster of bristles on the cucullus.

Distribution. New Guinea: West Papua ( Indonesia) and New Britain ( Papua New Guinea).

Figures

FIGURES 4−9. Labial palpi of Cimeliomorpha spp. 4. C. cymbalora, female (Thailand, KKIC, np2895). 5. C. novarana, female (Thailand, KKIC, np5789). 6. C. egregiana, male (Amboyna, Doherty, 1892 No.41343, NHMUK). 7. C. jarujini, male (Thailand, holotype). 8. C. perspinosa, male (West Papua, holotype). 9. C. inflata, male (Woodlark Id, holotype).

FIGURES 16−23. Adults of egregiana group (scale bars = 2 mm). 16. C. egregiana, male (Amboyna, Doherty, 1892 No.41343, NHMUK). 17. C. egregiana, female (Amboyna, Doherty, 1892 No.41344, NHMUK). 18. C. jarujini, male holotype (Thailand). 19. C. jarujini, female paratype (Thailand, KKIC, np1852). 20. C. perspinosa, male holotype (West Papua). 21. C. perspinosa, female paratype (Papua New Guinea, NHMUK). 22. C. inflata, male holotype (Woodlark Id). 23. C. inflata, female paratype (Woodlark Id, ANIC).

FIGURES 27−34. Male genitalia of egregiana group. 27. C. egregiana (Amboyna, B.M. genitalia slide no. 33120). 28. C. egregiana, spines (right valva). 29. C. jarujini (Thailand, holotype). 30. C. jarujini, spines (left valva). 31. C. perspinosa (West Papua, holotype). 32. C. perspinosa, spines (left valva). 33. C. inflata (Woodlark, holotype). 34. C. inflata, spines (left valva) (Paratype, B.M. genitalia slide no. 33131).

FIGURES 35−40. Socii of Cimeliomorpha spp. 35. C. cymbalora (Thailand, KKIC, genitalia slide NP1936). 36. C. novarana (Thailand, KKIC, genitalia slide NP2001). 37. C. egregiana (Amboyna, B.M. genitalia slide no. 33120). 38. C. jarujini (Thailand, holotype). 39. C. perspinosa (West Papua, holotype). 40. C. inflata (Woodlark, holotype)

FIGURES 55−58. Cimeliomorpha perspinosa (Papua New Guinea, B.M. genitalia slide no. 33122). 55. Female genitalia. 56. Ostium. 57−58. Signa.