Hydrochus bernardi Makhan

Perkins, Philip D., 2021, Taxonomy of Central American water beetles in the genus Hydrochus Leach, 1817 (Coleoptera: Hydrochidae), Zootaxa 4974 (3), pp. 459-503 : 473

Hydrochus bernardi Makhan

Figs. 1 , 20 , 41

Hydrochus bernardi Makhan 1994: 42 .

Originally described from Mexico. The following are new locality data records from Costa Rica and Mexico.

Material examined (44): Costa Rica: 16 m. S. La Cruz , 13.vii.1965, leg. Paul J. Spangler (21 NMNH) . Mexico: Chiapas, Mpio, Ángel Albino Corzo, Jaltenango , 16-Jan-2012, 15°52’15”N, 92°43’26” W, Trampa Luz, D. Reynoso col. (1 IEXA) ; Guerrero, 3.7 mi. E. Marquelia, 10.vii.1974, leg. Clark, Murray, Ashe, Schaffner (1 TAMU) ; 7 KM DE IXTAPA-PTO. LA SALITRERA, 28.vi.1986, leg. J. Bueno, S. Santiago (2 UNAM) ; Nayarit, Acaponeta , 15.viii.1960, leg. E. S. Ross, D. C. Rentz, P. H. Arnaud Jr. (1 CAS) ; San Blas , 17-21.ix.1953, leg. B. Malkin (1 CAS) ; Sinaloa, 7 mi. N. Rosario, 24.vii.1963, leg. Paul J. Spangler (17 NMNH) .

Differential Diagnosis. Among Central American species, recognized by the combination of moderate size (ca. 3.60 mm), the moderately densely punctate pronotum, and the male genitalia ( Fig. 20 ; described below). This species has a generalized Hydrochus habitus ( Fig. 20 ); reliable determinations will require dissection of males.

Male genitalia ( Fig. 20 ). General characters: paramere tips sagittate, plsa moderately prominent, narrower at very tip end in lateral view than in dorsal/ventral views, in dorsal/ventral views with medial margins at apex arcuate toward midline, then arcuate away from midline; aedeagus in dorsal/ventral views lanceolate, moderately wide at agw, at ca. midlength; in lateral view slightly narrower, without emargination where adbl and shaft meet; lateral margins of aedeagus sclerotized; basal piece slightly shorter than parameres, in dorsal/ventral views tapering from distal to proximal, in lateral view dorsal surface arcuate, ventral surfacsse straight, orifice strongly sclerotized.

Dorsal surface: pdmm widely separated, overlapping aedeagus; adbl moderately large; adtl subapical.

Ventral surface: pvmm approximating lateral margin of aedeagus; avtl not apparent; pseudoparameres cylindrical in cross section, ca. ½ as long as parameres.

Remarks. I have examined the holotype, including the male genitalia, of H. bernardi (ZMHB) ( Fig. 20 ).

Figures

FIGURE 1. Dorsal habitus of Central American Hydrochus species.

FIGURES 19–20. Habitus and male genitalia of holotypes. 19. Hydrochus elineae Makhan, non-type. 20. Hydrochus bernardi Makhan, holotype.

FIGURES 40–42. Geographical distribution maps.