Cacostola galenae Wappes and Santos-Silva, 2019

Wappes, James E. & Santos-Silva, Antonio, 2019, New species and taxonomic notes for Cacostola Fairmaire and Germain, 1859 (Coleoptera: Cerambycidae: Lamiinae: Onciderini), Insecta Mundi 741 (741), pp. 1-20 : 4-5

Cacostola galenae Wappes and Santos-Silva , new species

( Fig. 11–14 )

Description. Female. Integument mostly dark brown; mouthparts yellowish-brown; gulamentum dark reddish-brown; scape, pedicel, and antennomere III dark reddish-brown; antennomere IV reddish-brown basally, gradually brown toward apex; antennomeres V, VII, and IX yellowish-brown on basal quarter, brown on remaining surface; antennomeres VI and VIII yellowish-brown on basal half, gradually brown on distal half toward apex; antennomere X yellowish-brown on basal third, brown on remaining surface; Antennomere XI yellowish-brown basally and distally, brownish on wide central area; elytra and legs with irregular dark reddish-brown areas.

Head. Finely, sparsely punctate; pubescence not obscuring integument, pale yellow centrally and close to eyes (this latter area very narrow), yellowish brown on remaining surface, with short white setae interspersed throughout. Vertex with sculpturing as on frons; with pale yellow and yellowish-white pubescence partially obscuring integument, except glabrous narrow area along median groove. Area behind eyes finely punctate (punctures slightly denser than on vertex); with yellowish-brown pubescence partially obscuring integument, with short white setae interspersed. Genae 0.55 times length of lower eye lobe; sculpturing as on frons, except smooth apex; with yellowish-white pubescence not obscuring integument, sparser toward frons, especially on wide central area, except distal area glabrous. Antennal tubercles in frontal view widely U-shaped, with small inner projection; sculpturing as on frons, except smooth apex; with yellowish-white and pale-yellow pubescence mixed, denser toward posterior area. Postclypeus finely, sparsely punctate close to frons, finer, sparser toward anteclypeus; with sparse yellowish-brown pubescence on wide central area (nearly glabrous close to anteclypeus), glabrous laterally; with long, erect brownish setae on sides of wide central area close to anteclypeus. Labrum with sparse yellowish-brown setae centrally, longer, directed forward laterally; with fringe of nearly golden setae on anterior margin. Mandibles with a few short white setae on outer surface. Distance between upper eye lobes 0.45 times length of scape; in frontal view, distance between lower eye lobes 0.82 times length of scape. Antennae 1.6 times elytral length, reaching elytral apex at middle of antennomere IX; antennal segments with yellowish-white pubescence not obscuring integument, appearing to be whiter on lighter areas; antennomeres III–VI with sparse, thick, dark setae ventrally (more abundant on III). Antennal formula (ratio) based on length of antennomere III: scape = 0.84; pedicel = 0.17; IV = 1.15; V = 1.06; VI = 0.92; VII = 0.81; VIII = 0.65; IX = 0.61; X = 0.61; XI = 0.61.

Thorax. Prothorax wider than long; sides unarmed, nearly parallel-sided at anterior 2/3, slightly convergent at posterior third. Pronotum coarsely, sparsely punctate; with slightly distinct, wide white pubescent band centrally (partially interspersed with yellowish pubescence), sides of posterior third with white pubescence, and remaining surface with yellowish-brown pubescence; with a few long, erect dark setae near posterolateral angles. Sides of prothorax coarsely, sparsely punctate; with white and yellow pubescence mixed, partially obscuring integument. Ventral surface of thorax with dense white and yellow pubescence mixed laterally, yellowish-white, slightly less dense centrally; narrowest area of prosternal process 0.4 times width of procoxal cavity; mesoventral process 0.65 times width of mesocoxal cavity. Scutellum with sparse yellowish-white pubescence. Elytra. Humeri rounded, very slightly projected; coarsely, abundantly punctate on basal third, gradually finer, sparser toward apex; with dense, wide white pubescent band dorsally, from humerus to apex, longitudinally divided with yellowish-brown pubescent band from slightly before middle to posterior seventh, narrowed distally; sutural area with narrow white pubescent band, more distinct on basal third; area between sutural and lateral pubescent bands with sparse yellowish-brown pubescence, with white setae interspersed, from base to about posterior third (narrowed toward its apex); remaining surface with irregular white and yellowish-brown pubescent areas. Legs. Femora with white pubescence not obscuring integument, dorsally slightly yellower toward apex; protibiae with white pubescence not obscuring integument, except dark yellowish-brown pubescence on ventral surface of posterior 2/3; mesotibiae with white pubescence not obscuring integument, with pale yellow pubescence interspersed, except dark yellowish-brown pubescence on dorsal half, apex of sides, and distal third ventrally; metatibiae with white pubescence not obscuring integument, except distal third of posterior half with yellowish pubescence.

Abdomen. Ventrites I–IV with dense yellowish-brown pubescence laterally, sparser centrally, these two areas separated on each side by oblique band with denser pubescence, yellowish-brown on ventrite I, gradually lighter toward IV; with central band, from base to about middle denser pubescent. Ventrite V with dense yellowish-white pubescence laterally, with white pubescence interspersed, and slightly sparser pale-yellow pubescence centrally; apex concave.

Dimensions (mm). Total length, 7.80; prothoracic length, 1.25; anterior prothoracic width, 1.40; posterior prothoracic width, 1.30; maximum prothoracic width, 1.45; humeral width, 1.90; elytral length, 6.00.

Type material. Holotype female from PANAMA, Panama: Bayano dst. (3 km S Ipiti), 24.V.1992, J.E. Wappes col. ( FSCA, formerly ACMT).

Remarks. Cacostola galenae sp. nov. resembles C. janzeni Chemsak and Linsley, 1986 (see photograph of the holotype at Bezark 2019a), but differs by the antennal tubercles not prominent, nor forming a broad V-shape from above (prominent and V-shaped in C. janzeni ), and by the elytral punctures distinctly sparser (denser in C. janzeni ). It is also similar to C. colombiana Martins and Galileo, 1999 (see photograph of the holotype at Bezark 2019a), differing by having antennal tubercles more separated from each other, and antennae slightly surpassing the elytral apex (distinctly longer in C. colombiana ). It is also similar to C. grisea Dillon and Dillon, 1946 , but differs by the antennal tubercles not being distinctly prominent (prominent in C. grisea ), and by the area between antennal tubercles not distinctly concave (noticeably concave in C. grisea ).

Etymology. Named to thank and recognize Galena Opitz, a delightful lady and spouse of Weston Opitz, who not only provides him with great technical assistance in his systematic research on Cleridae but also welcomes his friends as hers, making them most welcome in their home.

Figures

Figures 11–19. Cacostola spp. 11–14) Cacostola galenae, holotype female. 11) Dorsal habitus. 12) Ventral habitus. 13) Lateral habitus. 14) Head, frontal view. 15–19) Cacostola opitzi, holotype male. 15) Dorsal habitus. 16) Ventral habitus. 17) Lateral habitus. 18) Head, frontal view. 19) Cacostola opitzi, paratype female, dorsal habitus.