Enicospilus javanus ( Szepligeti , 1910)

Shimizu, So, 2020, The Nepalese species of the genus Enicospilus Stephens, 1835 (Hymenoptera, Ichneumonidae, Ophioninae): a preliminary revision and identification key to species, Deutsche Entomologische Zeitschrift 67 (1), pp. 69-126 : 69

Enicospilus javanus ( Szepligeti, 1910) Fig. 12

Henicospilus javanus Szépligeti 1910: 93; holotype ♀, Java, TM.

Enicospilus fulacorensis Brues 1918: 117; holotype ♀, Solomon Island, MCZ; synonymised by Gauld and Mitchell (1981: 260).

Enicospilus gephyrus Chiu 1954: 32; holotype ♀, Japan, TARI, examined; synonymised by Gauld and Mitchell (1981: 260).

Enicospilus (Bicorniata) diurnus Nikam 1975: 193, 194; holotype ♀, India, MUC; synonymised by Gauld and Mitchell (1981: 260).

Material examined.

44♀♀ 4♂♂: Nepal (5♀♀ 2♂♂), Brunei (30♀♀ 2♂♂) India (2♀♀), Papua New Guinea (4♀♀), Singapore (1♀), Sri Lanka (2♀♀) .

Non-type series: 2♀♀ 1♂, Kakani (2,000 m), Nepal, VIII.1982, M.G. Allen leg. (LT) ; 3♀♀, Kathmandu (1,350 m), Nepal, VII.1983, M.G. Allen leg. (LT) (Fig. 12 ) ; 1♂, Pokhara , Nepal, VIII.1982, M.G. Allen leg. (LT) ; 2♀♀ 1♂, Gn. Pagon (1,700 m), U. Temburong, Brunei, IV.1981, I.D. Gauld leg. ; 24♀♀, Bukit Retak (1,500 m), U. Temburong, Brunei, IV.1981, I.D. Gauld leg. ; 1♂, Montane forest (1,618 m), Bukit Retak, Brunei, IX.1979, I.D. Gauld leg. ; 2♀♀, Pagon Ridge , Pagon, Brunei, II.1982, I.D. Gauld leg. ; 2♀♀, 1′ forest (500 m), U. Temburong, Brunei, IV.1981, I.D. Gauld leg. ; 1♀, Thekkadi , Periyar Dam, Travancore, India, 6-10.V.1937 ; 1♀, Andhra Pradesh, Patancheru , India, XII.1980, Bhatnagar leg. (LT) ; 1♀, Wau (1,200 m), Morobe Dist., Papua New Guinea, X.1979, I.D. Gauld leg. ; 3♀♀, Mt Lawes (400 m), Port Moresby, Papua New Guinea, 5.III-12.V.1963, W.W. Brandt leg. ; 1♀, Singapore, 1901, H.N. Kidley leg. ; 1♀, Peradeniya , Sri Lanka, 25.VII.1919, N.K. Jardine leg. ; 1♀, Matale (1,500 m), Sri Lanka, 10.V.1919, N.K. Jardine leg. (all NHMUK).

Distribution.

Australasian, Eastern Palaearctic, and Oriental regions ( Yu et al. 2016). Gauld and Mitchell (1981) recorded this species from Nepal.

Diagnosis.

Head (Fig. 12B-D ): GOI = 2.7-3.2; lower face 0.7-0.8 × as wide as high; clypeus moderately convex in profile, its lower margin blunt; mandible moderately twisted by 15-30°, moderately long, evenly tapered, its outer surface without a diagonal structure; upper mandibular tooth 1.2-1.5 × as long as lower one; posterior ocellus almost touching eye; antenna with 55-62 flagellomeres and 20th flagellomere 2.0-2.5 × as long as wide.

Mesosoma (Fig. 12E ): mesopleuron dorsally punctate to longitudinally striate and ventrally longitudinally striate; scutellum with lateral longitudinal carinae reaching posterior end and convergent posteriorly; metapleuron striate; propodeum declivous, its posterior area moderately reticulate, outer margin of propodeal spiracle joining pleural carina by a ridge.

Wings (Fig. 12F ): fore wing with AI = 1.1-1.8, CI = 0.2-0.5, ICI = 0.4-0.6, SDI = 1.0-1.1; fore wing vein 1m-cu&M evenly curved, 2r&RS almost straight; fenestra and sclerites of discosubmarginal cell of fore wing as in Figure 12F ; fenestra of fore wing not very long and its anterodistal corner distinctly separated from proximal end of vein RS; proximal sclerite not triangular, confluent with distal one, at least anteriorly very strongly pigmented; central sclerite absent; distal sclerite more or less strong and rather thick; proximal corner of marginal cell of fore wing more or less uniformly setose; vein 1cu-a subinterstitial to antefurcal to M&RS by less than 0.1 × 1cu-a length.

Colour (Fig. 12 ): body including interocellar area entirely testaceous; wings hyaline to slightly infumate.

Differential diagnosis.

Enicospilus javanus is distinctive and one of the most easily distinguishable species on account of the proximally extended fore wing fenestra and the shape of the sclerites (cf. Fig. 12F ).

Figures

Figure 12. Enicospilus javanus (Szepligeti, 1910), ♀. A. Habitus; B. Head, frontal view; C. Head, lateral view; D. Head, dorsal view; E. Mesosoma, lateral view; F. Central part of fore wing.