Liroetis modestus ( Weise, 1922 ) Bezděk, 2021

Bezděk, Jan, 2021, Redefinition of Liroetis, with descriptions of two new species and an annotated list of species (Coleoptera: Chrysomelidae: Galerucinae), Acta Entomologica Musei Nationalis Pragae 61 (2), pp. 529-614 : 578-580

Liroetis modestus ( Weise, 1922) comb. nov.

( Figs 192 , 204 , 215 , 232–238 )

Siemssenius modestus Weise, 1922: 73 (original description).

Siemssenius modestus : Wൾංඌൾ (1924): 131 (catalogue); Gඋൾඌඌංඍඍ & Kංආඈඍඈ (1963): 555 (noted); Wංඅർඈඑ (1971): 60 (catalogue); WൺඇǤ & YൺඇǤ (1998): 91 (noted); YൺඇǤ (2002): 638 (noted); ZIJൺඇǤ et al. (2008b): 127 (key); Bൾൾඇൾඇ (2010): 488 (catalogue); YൺඇǤ et al. (2015): 251 (key), 251 (noted).

Type locality. ‘Fokien [= China: Fujian]’.

Type material examined. Sඒඇඍඒඉൾඌ: 1 ♀ ( Figs 232–236 ), ‘ China / Prov. Fokien / G. Siemssen vend. / 14.11.1903 [w, p] // J. Weise determ. / 1919. [w, h] // Siemssenius / modestus / Cotype! [w, h] // Siemssenius / modestus / Ws [r, h]’ ( USNM); 1 ♀, ‘ China / Prov. Fokien / G. Siemssen vend. / 14.11.1903 [w, p] // J. Weise determ. / 1919. [w, h]’ ( USNM).

Additional material examined. CHINA: Fඎඃංൺඇ: Wuyishan Mts.NNR, Taoyuanyu valley, 27°43.7′N 117°42.8′E, 575 m, 1 J 1 ♀, J. Hájek,

D. Král, J. Růžička & L.Sekerka leg.( NMPC). ZΗൾඃංൺඇǤ: Tienmu Shan, 1 J, E. Reitter leg. ( NHMB).

Diagnosis. Colouration. Body pale brown except for darkened apices of mandibles, black antennae and legs with black tibiae, tarsi and apices of femora. Male from Zhejiang have paler basal and apical antennomeres and basal parts of tibiae.

Body length. J: 10.7 mm, ♀♀: 9.8–12.0 mm.

Male ( Fig. 237 ). Antennae 0.72 times as long as body. Pronotum 1.77 times as wide as long, covered with fine punctures, anterior margin moderately concave, unbordered, posterior margin thinly bordered, lateral margins parallel in posterior half, rounded and convergent anteriorly, with wider border. Posterior margin of abdominal ventrite IV with two transverse triangular processes separated by U-shaped incision. Longitudinal impression on last abdominal ventrite very wide and deep, constricted subbasally. Pygidium with elevated keel in posterior half. Metatibial spur wide, flat, with oblique apex.

Aedeagus ( Fig. 192 ). Median lobe of aedeagus 3.77 times as long as wide; apical tenth narrower, divergent apically, basal three quarters subparallel. Lateral view: median lobe bisinuate, apical quarter gradually bent; lateral elevation triangular, placed in anterior sixth of aedeagus length. Dorsal process long, 5.00 times as long as wide, 0.89 times as long as median lobe; basal third narrow, parallel, apical two thirds wider, lanceolate, with sharp apex. Lateral view: dorsal process wide, subparallel in apical and basal halves, bent in middle part, apex sharp, slightly bent downwards.

Female ( Fig. 238 ). Metatibial spur absent. Pygidium with blunt median keel. Sternite VIII subhexagonal, posterior margin with small wide triangular incision followed by short keel, surface laterally with two elevated plates, long setae accumulated along posterior margins of those plates; tignum 0.50 times as long as sternite VIII ( Fig. 215 ). Spermatheca with slightly indicated nodulus, cornu C-shaped, gradually narrowed towards sharp apex, spermathecal duct relatively long and straight ( Fig. 204 ).

Differential diagnosis. Liroetis modestus is very similar to L. fulvipennis . Both species can be distinguished by the colouration of legs (bicolour in L. modestus , black in L. fulvipennis ) and by dorsal process of aedeagus widest in anterior third in L. modestus but widest in the middle in L. fulvipennis ( Figs 187 , 192 ).

Distribution. China: Fujian (Wൾංඌൾ 1922, present paper), Zhejiang (present paper).

Figures

Figs 189–192. Aedeagus of Liroetis, dorsal and lateral views. 189 – L. jungchani (Lee, 2016); 190 – L. liui (Lee, 2016); 191 – L. metallipennis (Chûjô, 1962); 192 – L. modestus (Weise, 1922). Scale 0.5 mm.

Figs 197–208. 197 – Aedeagus of Liroetis yuae (Lee, 2016), dorsal and lateral views. 198–208 – Spermatheca. 198 – L. cheni (Lee, 2016); 199 – L. fulvipennis Jacoby, 1890; 200 – L. jeanvoinei (Laboissière, 1929); 201 – L. jungchani (Lee, 2016); 202 – L. liui (Lee, 2016); 203 – L. metallipennis (Chûjô, 1962); 204 – L. modestus (Weise, 1922); 205 – L. nigriceps (Laboissière, 1929); 206 – L. rufipennis (Chûjô, 1962); 207 – L. tsoui (Lee, 2016); 208 – L. yuae (Lee, 2016). Figs 198, 201–203, 207–208 reproduced from Lൾൾ (2016). Scales: 0.5 mm for Fig. 197, 0.25 mm for Figs 198–208.

Figs 209–219.Sternite VIII.209 – L. cheni (Lee, 2016); 210 – L. fulvipennis Jacoby,1890; 211 – L.jeanvoinei (Laboissière, 1929); 212 – L. jungchani (Lee, 2016); 213 – L. liui (Lee, 2016); 214 – L. metallipennis (Chûjô, 1962); 215 – L. modestus (Weise, 1922); 216 – L. nigriceps (Laboissière, 1929); 217 – L. rufipennis (Chûjô, 1962); 218 – L. tsoui (Lee, 2016); 219 – L. yuae (Lee, 2016). Figs 209, 212–214, 218–219 reproduced from Lൾൾ (2016). Scale 0.5 mm.

Figs 232–238. Liroetis modestus (Weise, 1922). 232–236 – Female, syntype (9.8 mm). 232 – dorsal view; 233 – lateral view; 234 – ventral view; 235 – head and pronotum; 236 – labels. 237–238 – Male (10.7 mm). 237 – dorsal view; 238 – lateral view.

Figs 185–188.Aedeagus of Liroetis, dorsal and lateral views.185 – L. cheni (Lee, 2016); 186 – L. elongatus (Kimoto, 1977); 187 – L. fulvipennis Jacoby, 1890; 188 – L. jeanvoinei (Laboissière, 1929). Scale 0.5 mm.