Cyrtodactylus bansocensis, Luu, Vinh Quang, Nguyen, Truong Quang, Le, Minh Duc, Bonkowski, Michael & Ziegler, Thomas, 2016

Luu, Vinh Quang, Nguyen, Truong Quang, Le, Minh Duc, Bonkowski, Michael & Ziegler, Thomas, 2016, A new species of karst-dwelling bent-toed gecko (Squamata: Gekkonidae) from Khammouane Province, central Laos, Zootaxa 4079 (1), pp. 87-102 : 91-98

Cyrtodactylus bansocensis sp. nov.

( Figs. 2–4 )

Holotype. VFU R.2015.20 , adult male, on karst cliff, above the entrance of Peopalam cave (17°27.101’N, 105°35.393’E, 195 m elevation), near Ban Soc Village, Bualapha District , Khammouane Province, central Laos, collected on 17 March 2015 by V. Q. Luu and K. Thanabuaosy.

Paratype. NUOL R- 2015.21, adult male, same collection data as the holotype.

Diagnosis. The new species can be distinguished from congeners by the following combination of characters: medium size, SVL reaching 74.0 mm; dorsal pattern with four light transverse bands between limb insertions; supranasals in contact; 14–15 irregular dorsal tubercle rows at midbody; lateral skin fold present without tubercles; 34–35 ventral scale rows between ventrolateral folds; 34 precloacal and femoral pores in a continuous row in the paratype and interrupted by four poreless scales in the holotype, enlarged femoral and precloacal scales present; 5– 7 postcloacal tubercles on each side; dorsal tubercles present at base of tail; subcaudal scales transversely enlarged, widely entire under tail surface.

Description of the holotype. Adult male, medium size (SVL 71.0 mm); body elongate (TrunkL/SVL 0.42); head distinct from neck, elongate (HL/SVL 0.28), relatively wide (HW/HL 0.64), depressed (HH/HL 0.41); loreal region concave; snout long (SE/HL 0.41), obtuse anteriorly, longer than diameter of orbit (OD/SE 0.64); scales on snout small, round, granular, bigger than those on frontal and parietal regions; eye large (OD/HL 0.26), pupils vertical; supraciliaries with spinuous scales posteriorly; ear oval-shaped, small (EarL/HL 0.11); rostral wider than high with a half median suture, in contact with first supralabial and nostril on each side; supranasals in broad contact; nostril opening oval, bordered by supranasal, rostral, first supralabial, and two enlarged postnasals; mental triangular, wider than long (ML/MW 0.86); two enlarged postmentals; supralabials 8/9; infralabials 8/8.

Dorsal scales granular; dorsal tubercles round, conical, present from occipital region to tail base, tubercles in 14 irregular rows at midbody, each surrounded by nine granular scales; lateral folds without tubercles; ventral scales smooth, round, largest posteriorly, in 35 longitudinal scales at midbody; gular region with homogeneous and smooth scales; 170 ventral scales from mental to cloacal slit; precloacal groove absent; enlarged femoral and cloacal scales present; total femoral and precloacal pores 34, on distal thigh, pore-bearing series interrupted by single scales lacking pores (1 + 1 + 29 + 3) ( Fig. 4 ).

Fore- and hindlimbs moderately slender (ForeL/SVL 0.17, CrusL/SVL 0.20); forelimbs lacking tubercles; dorsal surface of thigh and shank with distinctly developed tubercles, the same size to those on flanks; digits webbed basally; lamellae under fourth finger 16/17; lamellae under fourth toe 18/19.

Tail longer than SVL (TaL 98.5 mm, TaL/SVL 1.39); postcloacal tubercles 6/7; caudal tubercles present at the dorsum of the first segment of tail, extending from body; subcaudal scales transversely enlarged, three to four times as wide as long at base of tail, flat, smooth.

Coloration in life. Head dorsally greyish brown with dark blotches; distinct brown nuchal loop, widened in the nape, with dark edge, U–shaped, stretching from posterior edge of orbit to nape; labials grey. Dorsum with four light transverse bands between limb insertions, edged darker anteriorly, dark spots within two enlarged bands at midbody; tubercles at midbody yellowish brown; dorsal surface of fore and hind limbs with grey reticulated markings; tail greyish brown dorsally with 13 light rings (2–3 times narrower than dark tail bands); ventral surface of head, body and limbs cream; ventral surface of tail grey.

Variation: Light bands at midbody of the paratype (NUOL R-2015.21) are less distinctly defined at the posterior edges. The paratype also has precloacal and femoral pores in a continuous row (versus series interrupted by 4 poreless scales in the holotype). For further morphological characters of the paratype see Table 3 .

Comparisons. We compared Cyrtodactylus bansocensis sp. nov. with other species of Cyrtodactylus from Laos and neighbouring countries in the mainland Indochina region ( Vietnam, Cambodia, and Thailand). Comparisons were based on examination of museum specimens (see Appendix) and data from taxonomic publications ( Luu et al. 2014; Nazarov et al. 2014; Nguyen et al. 2014; Panitvong et al. 2014; Pauwels et al. 2014; Pauwels & Sumontha 2014; Schneider et al. 2014; Sumontha et al. 2015; Nguyen et al. 2015; Luu et al. 2015; Luu et al. 2016) (see Table 4).

Cyrtodactylus bansocensis sp. nov. has enlarged subcaudal scales and thus differs from the following species which lack this character state: C. bidoupimontis Nazarov, Poyarkov, Orlov, Phung, Nguyen, Hoang & Ziegler , C. bobrovi Nguyen, Le, Pham, Ngo, Hoang, Pham & Ziegler , C. buchardi David, Teynié & Ohler , C. bugiamapensis Nazarov, Poyarkov, Orlov, Phung, Nguyen, Hoang & Ziegler , C. cattienensis Geissler, Nazarov, Orlov, Böhme, Phung, Nguyen & Ziegler , C. cryptus Heidrich, Rösler, Vu, Böhme & Ziegler , C. cucdongensis Schneider, Phung, Le, Nguyen & Ziegler , C. huynhi Ngo & Bauer , C. interdigitalis Ulber , C. irregularis (Smith) , C. otai Nguyen, Le, Pham, Ngo, Hoang, Pham & Ziegler , C. phuocbinhensis Nguyen, Le, Tran, Orlov, Lathrop, Macculloch, Le, Jin, Nguyen, Nguyen, Hoang, Che, Murphy & Zhang , C. pseudoquadrivirgatus Rösler, Vu, Nguyen, Ngo & Ziegler , C. quadrivirgatus Taylor , C. ranongensis Sumontha, Pauwels, Panitvong, Kunya & Grismer , C. taynguyenensis Nguyen, Le, Tran, Orlov, Lathrop, Macculloch, Le, Jin, Nguyen, Nguyen, Hoang, Che, Murphy & Zhang , C. thuongae Phung , van Schingen, Ziegler & Nguyen, C. vilaphongi Schneider, Nguyen, Le, Nophaseud, Bonkowski & Ziegler , and C. ziegleri Nazarov, Orlov, Nguyen & Ho.

The new species has femoral and precloacal pores in males, which are absent in the following species: C. angularis (Smith) , C. badenensis Nguyen, Orlov & Darevsky , C. chauquangensis Hoang, Orlov, Ananjeva, Johns, Hoang & Dau , C. cucphuongensis Ngo & Chan , C. eisenmanae Ngo , C. grismeri Ngo , C. martini Ngo, C.

nigriocularis Nguyen, Orlov & Darevsky , C. oldhami (Theobald) , C. pageli Schneider, Nguyen, Schmitz, Kingsada, Auer & Ziegler , C. paradoxus (Darevsky & Szczerbak) , C. puhuensis Nguyen, Yang, Le, Nguyen, Orlov, Hoang, Nguyen, Jin, Rao, Hoang, Che, Murphy & Zhang , C. saiyok Panitvong, Sumontha, Tunprasert & Pauwels , C. samroiyot Pauwels & Sumontha , C. sanook Pauwels, Sumontha, Latinne & Grismer , C. spelaeus Nazarov, Poyarkov, Orlov, Nguyen, Milto, Martynov, Konstantinov & Chulisov , C. sumonthai Bauer, Pauwels & Chanhome , C. wayakonei Nguyen, Kingsada, Rösler, Auer & Ziegler , and C. thirakhupti Pauwels, Bauer, Sumontha & Chanhome.

The new species has 34 femoral and precloacal pores in males and thus differs from the following species which have distinctly fewer femoral and precloacal pores: C. auribalteatus Sumontha, Panitvong & Deein (10–11), C. bichnganae Ngo & Grismer (28), C. brevipalmatus (Smith) (22), C. caovansungi Orlov, Nguyen, Nazarov, Ananjeva & Nguyen (15), C. dumnuii Bauer, Kunya, Sumontha, Niyomwan, Pauwels, Chanhome & Kunya (19), C. erythrops Bauer, Kunya, Sumontha, Niyomwan, Panitvong, Pauwels, Chanhome & Kunya (28), C. huongsonensis Luu, Nguyen, Do & Ziegler (21–23), C. intermedius (Smith) (8–10), C. khelangensis Pauwels, Sumontha, Panitvong & Varaguttanonda (14–18), C. kingsadai Ziegler, Phung, Le & Nguyen (7–16), C. roesleri Ziegler, Nazarov, Orlov, Nguyen, Vu, Dang, Dinh & Schmitz (20–28), C. takouensis Ngo & Bauer (3–6), C. tigroides Bauer, Sumontha & Pauwels (21), and C. yangbayensis Ngo & Chan (6–10).

The new species has 34 femoral and precloacal pores in males and thus differs from the following species which have higher counts of femoral and precloacal pores: C. darevskii (38–44), C. jaegeri (44), C. jarujini Ulber (52–54), and C. multiporus (58–60).

The new species differs from C. chanhomeae Bauer, Sumontha & Pauwels by having fewer ventral scales (30–35 versus 36–38), fewer dorsal tubercles (14–15 versus 16–18), brown nuchal loop with dark edge (versus yellow edge), and white-grey body bands between limb insertions (versus yellow); from C. lomyenensis Ngo & Pauwels by having fewer dorsal tubercle rows (14–15 versus 20–24), fewer supralabials and infralabials (8–10 versus 13–14; 8 versus 11, respectively), and fewer femoral and precloacal pores in males (34 versus 39–40); from C. phongnhakebangensis Ziegler, Rösler, Herrmann & Vu by its smaller size (maximum SVL 74.0 mm versus 96.3 mm), the absence of tubercles on lateral folds (versus present), enlarged nuchal loop in neck region absent (versus present), four light transverse bands between limb insertions (versus 2–3), and tail with light rings (versus bands); and from C. teyniei David, Nguyen, Schneider & Ziegler by its smaller size (maximum SVL 74.0 mm versus 89.9 mm), having fewer ventral scale rows (34–35 versus 38), the presence of a nuchal band (versus being absent), and having a banded dorsal pattern (versus blotched).

Τaxa Cyrtodactylus bansocensis s. SvL ΤaL (mm) (mm).. ‾ 4.. ‾. v 4 EFS FP s s PP PP LD4 (in males) (in females) 4 unknοwn ‾ LΤ4 Cοlοr pattern Enlarged οf dοrsum subcaudals!" # # s C. rufford 68.3‾72.5 94.5‾96.8 27‾29 present present 42‾43 unknοwn 19‾20 18‾19 banded present C. soudthichaki 69.2‾70.0 95.1*‾95.2 32‾33 present present 29 absent 16-18 18 banded present C. angularis 80.0‾92.0 92‾95.2 40‾45 present absent 3 3 18 ‾19 18‾19 banded present C. astrum 46.4‾108.3 99.0*‾109.0* 31‾46 ‾ present 31‾38 ‾ ‾ (FP+PP) 20‾24 banded present C. auribalteatus 82.8‾98.1 106.5‾138.7 38‾40 5‾7 4‾5 (in 6 absent ‾ 18‾21 banded present C. badenensis 59.3‾74.1 58.6‾82.4 25‾29 absent absent 0 0 ‾ 18‾22 banded present C. bichnganae 95.3‾99.9 96.3‾115.6 30‾31 11‾13 18 10 8 18‾20 16‾20 banded present C. bidoupimontis 74.0‾86.3 75.0‾86.0 38‾43 6‾8 absent 4‾6 0 15‾20 18‾23 banded absent C. bobrovi 75.2‾96.4 80.8‾90.3 40‾45 0 0 5 0 19‾21 21‾22 banded absent C. brevipalmatus 64.0‾72.0 77.0 35‾44 present present 6+9+7 6+9+7 ‾ ‾ blοtched present C. bugiamapensis 58.6‾76.8 65.3‾83.0 36‾46 6‾10 absent 7‾8 0‾7 15‾17 17‾20 blοtched absent C. buchardi 60.0‾65.0 46.0‾54.0 30 absent absent 9 0 14 12 blοtched absent C. caovansungi 90.4‾94.0 120.0 38‾44 8 6 9 0 22 23‾25 banded present C. cattienensis 43.5‾ 69.0 51.0‾64.7 28‾42 3‾8 absent 6‾8 0 12‾16 14‾19 banded absent C. chanhomeae 69.9‾78.8 74.4‾74.7 36‾38 present present 32 34 18‾20 21‾23 banded present C. chauquangensis 90.9‾99.3 97.0‾108.3 36‾38 absent absent 6 7 16‾18 19‾23 banded present C. cryptus 62.5‾90.8 63.5‾88.4 47‾50 absent absent 9‾11 0 18‾19 20‾23 banded absent C. cucdongensis 55.8‾65.9 22.1‾27.8 35‾44 present absent 5‾6 4‾6 8‾11 15‾20 banded absent C. cucphuongensis 96.0 79.3* 42 14 absent 0 ‾ 21 24 banded present C. darevskii 84.6‾100.0 95.0‾113.0 38‾46 present present 38‾44 24‾34 17‾20 (FP+PP) 18‾22 banded present

(in males) (in males) (FP+PP) (FP+PP) males) (FPl+PP+FPr) (FPl+PP+FPr) (FP+PP) (FP+PP)

Τaxa SvL (mm) ΤaL (mm) v EFS FP PP (in males) PP (in females) LD4 LΤ4 Cοlοr pattern οf dοrsum Enlarged subcaudals C. dumnuii 76.2‾84.2 100.2* 40 present present in males⁄absent 6+5‾6+6‾7 (FPl+PP+FPr) 0‾7 16 19 banded present C. eisenmanae 76.8‾89.2 91.0‾103.8 44‾45 4‾6 absent 0 0 18‾20 17‾18 banded present C. erythrops 78.4 83.0* 28 present present 10+9+9 ‾ 16 20 blοtched present C. grismeri 68.3‾95.0 111.3‾115.1 33‾38 absent ‾ 0 0 16‾18 16‾19 banded present C. huongsonensis 73.4‾89.8 90.5 41‾48 7‾9 15‾17 6 8 17‾19 20‾23 banded present C. huynhi 54.8‾79.8 61.5‾78.6 43‾46 3‾5 3‾8 7‾9 0‾8 14‾17 17‾21 banded absent C. interdigitalis 59.0‾80.0 71.0‾90.0 37‾42 present 16‾18 14 0 17‾22 16‾20 banded absent C. intermedius 61.0‾85.0 80.0‾110.0 40‾50 6‾10 ‾ 8‾10 ‾ 20 22 banded present C. irregularis 72.0‾86.0 66.0‾74.0 38‾45 7‾8 ‾ 5‾7 0‾6 15‾16 18‾19 blοtched absent C. jaegeri 60.0‾68.5 82.4‾83.4 31‾32 17‾19 present 44 (FP+PP) 21 17‾19 20‾23 banded present C. jarujini 85.0‾90.0 105.0‾116.0 32‾38 present present 52-54 (PP+FP) 0 15‾17 18‾19 blοtched present C. khammouanensis 70.8‾73 83.0‾95.0 32‾38 present present 40‾44 (PP+FP) 0-17 18‾20 20‾23 banded present C. khelangensis 72.8‾95.3 max. 96.0* 32‾35 present present 6+2‾5+6‾7 2+6+1 18 22 banded present C. kingsadai 83.0‾94.0 max. 117.0 39‾46 9‾12 0‾7 7‾9 4‾8 19‾21 21‾25 banded present C. lekaguli 80.5‾103.5 115.0‾125.0 31‾43 present present 30‾36 0 ‾ 20‾25 banded present C. lomyenensis 57.7‾71.2 72.2‾86.1 35‾36 17‾18 present 39‾40 32 16‾19 19‾23 banded present C. martini 64.4‾96.2 76.0‾101.2 39‾43 14‾18 absent 4 0 19‾23 22‾24 banded present C. multiporus 81.0‾98.0 97.0‾105.0 30‾38 present present 58‾60 (PP+FP) 0 18‾20 18‾22 banded present C. nigriocularis 82.7‾107.5 70.6‾121 42‾49 absent absent 0‾2 0 ‾ 17‾21 unifοrmly brοwn present C. oldhami 63.0‾68.0 69*‾70* 34‾38 present absent 1‾4 ‾ ‾ ‾ striped and present C. otai 85.2‾90.6 89.7‾97.6 38‾43 absent absent 7‾8 0 16‾19 19‾22 banded absent C. pageli 76.2‾81.8 85.4*‾113.2* 41‾44 absent absent 4 4 19‾23 19‾23 banded present

in females

(FPl+PP+FPr)

(FPl+PP+FPr) (FPl+PP+FPr)

(PP+FP) (PP+FP) (PP+FP)

spοtted

……continued on the next page

Τaxa SvL ΤaL v EFS FP PP PP LD4 LΤ4 Cοlοr pattern Enlarged C. paradoxus 52.0‾84.0 80.8‾111.0 32‾40 present absent 0‾4 0 15‾18 17‾23 banded present C. phongnhakebangensis C. phuocbinhensis 78.5‾96.3 46.0‾60.4 98.0‾110.0 76.1 32‾42 43‾47 present 5 present absent 32‾42 (PP + FP) 7 0‾41 (PP + FP) 0 15‾20 16‾21 18‾26 17‾19 banded blοtched present absent C. pseudoquadrivirgatus 48.6‾83.3 55.7‾82.3 41‾57 absent absent 5‾9 5‾10 15‾21 16‾25 blοtched absent C. puhuensis 79.2 82.59 36 present absent 5 ‾ 18 23 banded present C. quadrivirgatus 39.0‾67.0 77.0 40 present absent 4 4 ‾ ‾ striped absent C. ranongensis 56.9‾59.6 66.0‾67.1 35‾40 present 0 0 0 17 18 blοtched absent C. roesleri C. saiyok 51.1‾75.3 56.7‾61.0 63.4‾101.0 66.7‾67.5 34‾40 23‾24 7‾10 present present absent 20‾28 (PP + FP) 5 17‾22 (PP + FP) ‾ 17‾19 ‾ 17‾21 16‾17 banded banded present present C. samroiyot 63.2‾66.9 78.8‾87.5 33‾34 present absent 7 6 18 19 banded present C. sanook 72.9‾79.5 104.2 27‾28 present absent 3‾4 absent ‾ 19‾20 banded present C. spelaeus 88.9‾91.0 max. 83* 36‾39 absent absent 8‾9 0 19‾20 22‾24 banded present C. sumonthai 61.5‾70.7 89.9‾94.0 33‾36 absent absent 2 0 16 18 banded present C. takouensis 74.7‾81.1 77.7‾91.0 39‾40 3‾5 0‾2 3‾4 0 16‾17 18‾20 banded present C. taynguyenensis 60‾85 66‾94 42‾49 absent absent 6 0 13‾18 17‾21 blοtched absent C. teyniei 89.9 ca. 110.0 38 23 absent unknοwn 13 17‾18 19‾20 blοtched present C. thuongae 57.3‾77.6 max. 78.1 29‾44 2‾5 0‾3 0‾1 0 14‾17 14‾20 blοtched absent C. wayakonei 72.0‾86.8 76.8‾89.0 31‾35 absent absent 6‾8 7 17‾18 19‾20 banded present C. thirakhupti 72.0‾79.6 99.1 37‾40 present absent absent absent 16 20 banded present C. tigroides C. vilaphongi 74.3‾83.2 60.9‾86.1 108.5‾117.0 61.2‾68.1 34 34‾36 present 0 present ‾ 6+8+7 (FPl+PP+FPr) ‾ 5+9+7 (FPl+PP+FPr) 0 18‾19 18‾19 20‾22 18‾20 banded banded present absent C. yangbayensis 78.5‾92.3 91.3‾109.1 39‾46 5‾16 0‾2 6‾8 0 16‾19 15‾17 banded present C. ziegleri 84.6‾93.0 95.0‾107.0 33‾39 8‾10 0‾6 5‾8 0‾8 16‾19 18‾21 banded absent

(mm) (mm) (in males) (in females) οf dοrsum subcaudals

Cyrtodactylus bansocensis sp. nov. is morphologically similar to C. rufford ( Luu et al. 2016) , C. khammouanensis , and C. soudthichaki in the dorsal colour pattern and/or numbers of femoral and precloacal pores. However, the new species can be distinguished from C. rufford by having more ventral scale rows (34–35 versus 27–29), fewer supralabials (8–10 versus 10–12), fewer infralabials (8 versus 9–11), fewer femoral and precloacal pores in males (34 versus 42–43), and more postcloacal tubercles on each side (5–7 versus 4–5); C. khammouanensis by having fewer dorsal tubercle rows (14–15 versus 16–21), having fewer supralabials and infralabials (8–10 versus 11–12; 8 versus 9–10, respectively), fewer femoral and precloacal pores in males (34 versus 40–44), and tail with light rings (versus light bands); and from C. soudthichaki by its larger size (SVL reaching 74.0 mm versus 70.0 mm), having fewer dorsal tubercle rows (14–15 versus 19–20), more femoral and precloacal pores in males (34 versus 29), slightly higher number of ventral scales (34–35 versus 32–33), more subdigital lamellae on fourth toe (18–21 versus 18), more postcloacal tubercles on each side (5–7 versus 4–5), and tail with light rings (versus light bands). For more details see Table 5 .

Distribution. Cyrtodactylus bansocensis sp. nov. is currently known only from the type locality in the karst forest near Ban Soc Village, Bualapha District, Khammouane Province, central Laos ( Fig. 5 ).

Etymology. We name this species after its type locality, Ban Soc limestone forest to underline the importance of this area (see also Ziegler et al. 2015) for biodiversity and nature conservation. We suggest as common names: Ki Chiem Ban Soc (Laotian), and Ban Soc Bent-toed Gecko (English).

Natural history. The type specimens of the new species were collected between 20:00 and 21:00, on karst outcrops above the entrance of Peopalam cave, at an elevation of 195 m a.s.l. The surrounding habitat was secondary forest dominated by the species of Ebenaceae, Arecaeae , Poaceae , Meliaceae , and Moraceae . The benttoed geckos were actively foraging on surfaces of karst outcrops. They were fast and difficult to approach and catch ( Fig. 6 ).

Figures

FIGURE 2. Dorsal view of the adult male holotype of Cyrtodactylus bansocensis sp. nov. (VFU R. 2015.20) in life. Photo: V. Q. Luu.

FIGURE 3. Details of the holotype of Cyrtodactylus bansocensis sp. nov. (VFU R. 2015.20) A) Dorsal head; B) ventral head; C) dorsal body; D) ventral body. Photos: V. Q. Luu.

FIGURE 4. Cloacal region of the holotype of Cyrtodactylus bansocensis sp. nov. (VFU R. 2015.20) (precloacal and femoral pores marked with x). Photo: V. Q. Luu.

FIGURE 5. Type locality (red circle) of Cyrtodactylus bansocensis in Khammouane Province, central Laos.

FIGURE 6. Habitat of Cyrtodactylus bansocensis. Photo: V. Q. Luu.

Tables

TABLE 3. Measurements (in mm) and morphological characters of the type series of Cyrtodactylus bansocensis sp. nov. (for other abbreviations see material and methods).

Character Sex	VFU R.2015.20 holotype male	NUOL R-2015.21 paratype male
SVL TaL	71.0 98.5	74.0 103.5
HH HL HW	8.3 20.2 13.0	7.5 21.0 13.3
OD SE EyeEar	5.3 8.3 5.7	5.5 8.7 5.3
EarL TrunkL ForeL	2.3 29.8 12.2	2.4 33.4 11.7
FemurL CrusL RW	15.7 14.1 3.1	16.7 15.7 2.8
RH MW ML	1.6 2.9 2.5	2.0 3.3 2.2
SL IL N	8/9 8/8 3/3	9/10 8/8 3/3
IN PM DTR	0 2 14	0 2 15
GST V SLB	9 35 170	9 34 158
SR FP+PP PAT	87 34 6/7	86 34 6/5
LD4 LT4	16/17 18/19	18/19 21/20

TABLE 5. Comparison of Cyrtodactylus bansocensis sp. nov. with other morphologically similar species of Cyrtodactylus (data obtained from Nazarov et al. 2014; Luu et al. 2015; Luu et al. 2016 and own data).

Character	Cyrtodactylus bansocensis sp. nov.	C. rufford	C. khammouanensis	C. soudthichaki
Number of specimens	2	3	4	3
Maximal snout–vent length (mm)	74.0	72.5	73.0	70.0
Ventral scales	34–35	27–29	32–38	32–33
Dorsal tubercle rows	14–15	14–16	16–21	19–20
Supralabials	8–10	10–12	11–12	10–11
Infralabials	8	9–11	9–10	8–9
Femoral and precloacal pores (in males)	34	42–43	40–44	29
Postcloacal tubercles	5–7	4–5	5–6	4–5
Transverse dorsal color pattern between limbs	light bands narrower than dark bands with dark spots in the mid- dorsal region	light thin sometimes irregular shaped bands as narrow as haft dark bands	light regular bands nearly as wide as dark bands	light bands wider than dark bands with dark blotches in the dorsolateral region
Tail color pattern	light rings	light rings	light bands	light bands