Amphineurus (Rhamphoneurus) anchoralis, Santos & Santos & Ribeiro, 2022

Santos, Daubian, Santos, Rodrigo dos Reis & Ribeiro, Guilherme Cunha, 2022, Systematics of Amphineurus (Rhamphoneurus Alexander) (Diptera: Tipuloidea: Limoniidae), Arthropod Systematics & amp; Phylogeny 80, pp. 439-494 : 439

3.3.3. Amphineurus (Rhamphoneurus) anchoralis sp. nov.

Material examined.

Holotype: ♂, Chile, Arauco, Nahuelbuta, Contulmo, Palo Botado [37°39′S 72°54′W], 12-II-1952, Peña (USNM) *. Paratypes: 1 ♂ and 1 [sex unknown], Chile, Arauco, Nahuelbuta [37°46′S 72°59′W], 2-II-1953, Peña (USNM); 1 [sex unknown], Chile, Chiloé Is., Rio Coluco [42°6′S 73°55′W], 30-I-1952, Peña (USNM) .

Etymology.

The word Amphineurus anchoralis is Latin for "of the anchor". The term refers to the shape of the projection of the male 9th tergite, which resembles an anchor.

Diagnosis.

The species is distinguished by the absence of markings near R1, and R2+3 shorter than R2. Furthermore, this species has a long vertical spur on the sheath of aedeagus, male tergite IX has a distinctive ridge projection, mesal lobes absent and the posterior margin of the male tergite IX is mostly unpigmented.

Description.

Wing length 6.15 mm, width 1.86 mm. - Coloration: General coloration yellowish brown. Rostrum and palpus dark brown. First segments of antenna pale, remaining segments dark yellow. Head brownish-black. Eyes black. Thorax yellowish-brown, pleura pale. Scutum without stripes. Halter pale with knobs light orange. Coxae and legs brownish. - Head (dorsal view Fig. 10B , ventral view Fig. 10A ): Rostrum bifurcated with short appendices; first palpal segment as long as terminal segment, second palpal segment dilated; scape thinner than pedicel, terminal flagellomere dilated, antenna with several long setae. - Thorax (lateral view Fig. 10E , dorsal view Fig. 10F ): a few setae on prescutum. Division line of laterotergite almost vertical; anatergite as tall as katatergite. Halter with dilated knob. Wing (Fig. 10D ) almost clear of markings, except marking covering R2; veins near fork of bM (Fig. 10C ) faded; M1+2 longer than basal deflection of M1; R2+3+4 at least twice as long as R2+3. - Male terminalia (Fig. 10G ): Male tergite IX with posterior margin with irregular ridges; median U-shaped notch with straight projection in middle; tip of projection also ridge shaped; projection and surrounding area darkened. Ventral branch of gonocoxite inclined laterally. Dorsal branch of gonocoxite half-length of ventral branch, atrophied and pointed medially; both branches setose. Lobe of gonostylus with pointed lobule, shaped similar to folded leaf. Clasper of gonostylus with differently shaped branches: lateral branch long, bent near middle; medial branch curved into stub. Mesal lobes absent. Sheath of aedeagus darkened on apical half, angulated, with long pointed projection perpendicular to remainder of sheath.

Remarks.

Specimens of this new species were previously identified by C.P. Alexander as A. (R.) sanus Alexander and A. (R.) insanus Alexander. This species resembles A. (R.) sanus but differs mainly in M1+2, shape of projection in male tergite IX, ventral branch of gonocoxite and spur of sheath of aedeagus.

Figures

Figure 10. Amphineurus (Rhamphoneurus) anchoralis sp. nov. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; k, knob of the halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; t 9, male tergite IX.